Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
The Nothosauria were much smaller reptiles than the plesiosaurs, none
of them perhaps exceeding the size of the smallest known plesiosaurs.
They were semiaquatic in habit, with many curious resemblances to
other semiaquatic reptiles of a later time known, as the dolichosaurs.
The neck is more or less elongated, having about twenty vertebrae in
the longest-necked forms; the body is moderately long, and broad,
and the tail is relatively short. The vertebrae and ribs are quite
like those of the plesiosaurs, that is, the vertebrae are gently
concave at each end, and the dorsal ribs are attached by a single
head to the transverse process high up on the arch; the cervical ribs
are double-headed, precisely like those of the older plesiosaurs,
one of the characters which insistently proves the relationships of
the two groups. The bones of the shoulders (Fig. 46) also have many
resemblances to the extraordinary ones of the plesiosaurs, though
they are much less specialized. There was no sternum; the coracoids
are large, though very much smaller than those of the plesiosaurs.
The collar-bones are large and strong, joining each other in front
of the coracoids and firmly united with the shoulder-blades at the
outer extremity. Four vertebrae are united to form a sacrum, and their
union with the hip bones (Fig. 47) was much firmer than was the
case with the plesiosaurs. The limbs are elongated, but it will be
observed in the figures (Fig. 48) that the radius and ulna, tibia and
fibula, that is, the bones of the forearm and of the leg proper, are
relatively very short as compared with the humerus and femur, a sure
indication of the beginning of aquatic habits. The toes and fingers
were doubtless webbed, and there was no increase in the numbers of
bones in the digits, so conspicuous in the plesiosaurs. The external
nostrils are large, but are not situated so far back near the eyes as
in the plesiosaurs. There is a large pineal opening in the top of the
skull, as in the plesiosaurs, but no sclerotic or bony plates have
been observed in the eyes. They had ventral ribs like those of the
plesiosaurs.
[Illustration: FIG. 48.—Legs of _Lariosaurus balsami_, an Upper
Triassic nothosaur: _h_, humerus; _r_, radius; _u_, ulna; _i_,
intermedium; _ue_, ulnare; _f_, femur; _fi_, fibula; _t_, tibia; _a_,
astragalus; _c_, calcaneum. (After Abel.)]
No impressions of scales or bony plates have ever been found with the
remains of the nothosaurs, and it is the belief that the skin was
bare. A good idea of their general appearance will be gained from the
accompanying restoration adapted from that of Professor Fraas (Fig. 44)
and the restoration of the less highly specialized _Lariosaurus_, made
from a very complete skeleton in the Frankfort museum (Fig. 49).
Public-domain text, read in full here on John Shaqi.
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