But in other woods, the ends of some cells which meet endwise
are absorbed, thus forming a continuous series of elements which
constitute an open tube. Such tubes are known as pores, or vessels,
or "tracheæ," and sometimes extend thru the whole stem. Besides this
marked difference between the porous and non-porous woods, the porous
woods are also distinguished by the fact that instead of being made
up, like the conifers of cells of practically only one kind, namely
tracheids, they are composed of several varieties of cells. Besides
the tracheae and tracheids already noted are such cells as "wood
fiber," "fibrous cells," and "parenchyma." Fig. 19. Wood fiber proper
has much thickened lignified walls and no pits, and its main function
is mechanical support. Fibrous cells are like the wood fibers except
that they retain their protoplasm. Parenchyma is composed of vertical
groups of short cells, the end ones of each group tapering to a point,
and each group originates from the transverse division of one
cambium cell. They are commonly grouped around the vessels (tracheæ).
Parenchyma constitutes the pith rays and other similar fibers, retains
its protoplasm, and becomes filled with starch in autumn.
[Illustration: Fig. 19. Isolated Fibers and Cells. _a_, four cells of
wood parenchyma; _b_, two cells from a pith ray; _c_, a single cell
or joint of a vessel, the openings, x, x, leading into its upper and
lower neighbors; _d_, tracheid; _e_, wood fiber proper. _After Roth._]
The most common type of structure among the broad-leaved trees
contains tracheæ, trachæids, woody fiber, fibrous cells and
parenchyma. Examples are poplars, birch, walnut, linden and locust. In
some, as ash, the tracheids are wanting; apple and maple have no woody
fiber, and oak and plum no fibrous cells.
This recital is enough to show that the wood of the broad-leaved trees
is much more complex in structure than that of the conifers. It is by
means of the number and distribution of these elements that particular
woods are identified microscopically. See p. 289.
[Illustration: Fig. 20.]
_Ring-porous woods._ Looking thru the microscope at a cross-section of
ash, a ring-porous wood, Fig. 20:
(1) The large round or oval pores or vessels grouped mostly in the
spring wood first attract attention. Smaller ones, but still quite
distinct, are to be seen scattered all thru the wood. It is by the
number and distribution of these pores that the different oak woods
are distinguished, those in white oak being smaller and more numerous,
while in red oak they are fewer and larger. It is evident that the
greater their share in the volume, the lighter in weight and the
weaker will be the wood. In a magnified cross-section of some woods,
as black locust, white elm and chestnut, see Chap. III, beautiful
patterns are to be seen composed of these pores. It is because of the
size of these pores and their great number that chestnut is so weak.
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