[Fibro-elastic Tissue.—We have already seen that when little cubes of
meat, &c., were placed on leaves, the muscles, areolar tissue, and
cartilage were completely dissolved, but the fibro-elastic tissue, even
the most delicate threads, were left without the least signs of having
been attacked. And it is well known that this tissue cannot be digested
by the gastric juice of animals.*
Mucin.—As this substance contains about 7 per cent. of nitrogen, I
expected that it would have excited the leaves greatly and been
digested by the secretion, but in this I was mistaken. From what is
stated in chemical works, it appears extremely doubtful whether mucin
can be prepared as a pure principle. That which I used (prepared by Dr.
Moore) was dry and hard. Particles moistened with water were placed on
four leaves, but after two days there was only a trace of inflection in
the immediately adjoining tentacles. These leaves were then tried with
bits of meat, and all four soon became strongly inflected. Some of the
dried mucin was then soaked in water for two days, and little cubes of
the proper size were placed on three leaves. After four days the
tentacles
* See, for instance, Schiff, ‘Phys. de la Digestion,’ 1867, tom. ii.,
p. 38. [page 123]
round the margins of the discs were a little inflected, and the
secretion collected on the disc was acid, but the exterior tentacles
were not affected. One leaf began to re-expand on the fourth day, and
all were fully re-expanded on the sixth. The glands which had been in
contact with the mucin were a little darkened. We may therefore
conclude that a small amount of some impurity of a moderately exciting
nature had been absorbed. That the mucin employed by me did contain
some soluble matter was proved by Dr. Sanderson, who on subjecting it
to artificial gastric juice found that in 1 hr. some was dissolved, but
only in the proportion of 23 to 100 of fibrin during the same time. The
cubes, though perhaps rather softer than those left in water for the
same time, retained their angles as sharp as ever. We may therefore
infer that the mucin itself was not dissolved or digested. Nor is it
digested by the gastric juice of living animals, and according to
Schiff* it is a layer of this substance which protects the coats of the
stomach from being corroded during digestion.