Haematin.—Some dark red granules, prepared from bullock’s blood, were
given me; these were found by Dr. Sanderson to be insoluble in water,
acids, and alcohol, so that they were probably haematin, together with
other bodies derived from the blood. Particles with little drops of
water were placed on four leaves, three of which were pretty closely
inflected in two days; the fourth only moderately so. On the third day
the glands in contact with the haematin were blackened, and some of the
tentacles seemed injured. After five days two leaves died, and the
third was dying; the fourth was beginning to re-expand, but many of its
glands were blackened and injured. It is therefore clear that matter
had been absorbed which was either actually poisonous or of too
stimulating a nature. The particles were much more softened than those
kept for the same time in water, but, judging by the eye, very little
reduced in bulk. Dr. Sanderson tried this substance with artificial
digestive fluid, in the manner described under globulin, and found that
whilst 1.31 of fibrin, only 0.456 of the haematin was dissolved in an
hour; but the dissolution by the secretion of even a less amount would
account for its action on Drosera. The residue left by the artificial
digestive fluid at first yielded nothing more to it during several
succeeding days.]
_Substances which are not Digested by the Secretion._
All the substances hitherto mentioned cause prolonged inflection of the
tentacles, and are either completely or at least partially dissolved by
the secretion. But there are many other substances, some of them
containing nitrogen, which are not in the least acted on by the
secretion, and do not induce inflection for a longer time than do
inorganic and insoluble objects. These unexciting and indigestible
substances are, as far as I have observed, epidermic productions (such
as bits of human nails, balls of hair, the quills of feathers),
fibro-elastic tissue, mucin, pepsin, urea, chitine, chlorophyll,
cellulose, gun-cotton, fat, oil, and starch. [page 122]
To these may be added dissolved sugar and gum, diluted alcohol, and
vegetable infusions not containing albumen, for none of these, as shown
in the last chapter, excite inflection. Now, it is a remarkable fact,
which affords additional and important evidence, that the ferment of
Drosera is closely similar to or identical with pepsin, that none of
these same substances are, as far as it is known, digested by the
gastric juice of animals, though some of them are acted on by the other
secretions of the alimentary canal. Nothing more need be said about
some of the above enumerated substances, excepting that they were
repeatedly tried on the leaves of Drosera, and were not in the least
affected by the secretion. About the others it will be advisable to
give my experiments.