If a tentacle is examined some hours after the gland has been excited
by repeated touches, or by an inorganic or organic particle placed on
it, or by the absorption of certain fluids, it presents a wholly
changed appearance. The cells, instead of being filled with homogeneous
purple fluid, now contain variously shaped masses of purple matter,
suspended in a colourless or almost colourless fluid. The change is so
conspicuous that it is visible through a weak lens, and even sometimes
by the naked eye; the tentacles now have a mottled appearance, so that
one thus affected can be picked out with ease from all the others. The
same result follows if the glands on the disc are irritated in any
manner, so that the exterior tentacles become inflected; for their
contents will then be found in an aggregated condition, although their
glands have not as yet touched any object. But aggregation may occur
independently of inflection, as we shall presently see. By whatever
cause the process may have been excited, it commences within the
glands, and then travels down the tentacles. It can be observed much
more distinctly in the upper cells of the pedicels than within the
glands, as these are somewhat opaque. Shortly after the tentacles have
re-expanded, the little masses of protoplasm are all redissolved, and
the purple fluid within the cells becomes as homogeneous and
transparent as it was at first. The process of redissolution travels
upwards from the bases of the tentacles to the glands, and therefore in
a reversed direction to that of aggregation. Tentacles in an aggregated
condition were shown to Prof. Huxley, Dr. Hooker, and Dr. Burdon
Sanderson, who observed the changes under the microscope, and were much
struck with the whole phenomenon. [page 40]
The little masses of aggregated matter are of the most diversified
shapes, often spherical or oval, sometimes much elongated, or quite
irregular with thread- or necklace-like or club-formed projections.
They consist of thick, apparently viscid matter, which in the exterior
tentacles is of a purplish, and in the short distal tentacles of a
greenish, colour. These little masses incessantly change their forms
and positions, being never at rest. A single mass will often separate
into two, which afterwards reunite. Their movements are rather slow,
and resemble those of Amoebae or of the white corpuscles of the blood.
We
FIG. 7. (Drosera rotundifolia.) Diagram of the same cell of a tentacle,
showing the various forms successively assumed by the aggregated masses
of protoplasm.