The Pisan fleet consisted of three squadrons. Of these, Admiral Andrea Morosini commanded the first; while the second was under Count Ugolino, who, in spite of his courage, was no trustworthy leader, on account of the devouring personal ambition urging him to subordinate the interests of the State to his own greed for power. The third was commanded by Andreotto Saracini. Oberto Doria, an officer of great courage and experience, was high admiral of the Genoese fleet. As it first hove in sight, this armada seemed no greater than that of the Pisans, but only because a reserve of thirty galleys, commanded by Benedetto Zaccaria, lay hidden behind Meloria--or, according to other accounts, behind Montenero--ready to join in the fight when required. Soon after midday the battle began, and raged for some hours without any decisive result. But when the two flag-ships met, both fleets closed in a general engagement. On either side vast numbers of combatants, killed, wounded, or stunned, were hurled into the sea. The waves were crimsoned with blood; drowning men clutched at oars to save their lives, but were relentlessly thrust under by the rowers' next strokes, owing to the impossibility of checking the manoeuvres in the thick of the fight, and at the most critical moment. Just then Benedetto Zaccaria, having been signalled for, hove in sight, full sail, and with sweeping oars, in time to decide the fate of the day. Seeing him draw near, the Pisans knew they were outnumbered, and their courage began to fail, although they continued the fight with undiminished ardour. As Zaccaria dashed in, he contrived to bring his galley alongside Doria's, so as to wedge Morosini, whose flag-ship was making a gallant defence. At the same time the galley bearing the Pisan standard was also surrounded by the foe. On all sides the sudden arrival of the reserve squadron had given fresh courage to the Genoese and diminished the hopes of Pisa. The struggle was now too unequal; nevertheless, both sides were unwilling to end it, for each bitter enemy was seeking to destroy not only the other's fleet, but the very life of the rival Republic.