For a thing to be a natural purpose in the _first_ place it is
requisite that its parts (as regards their being and their form) are
only possible through their reference to the whole. For the thing
itself is a purpose and so is comprehended under a concept or an Idea
which must determine _a priori_ all that is to be contained in it.
But so far as a thing is only thought as possible in this way, it is
a mere work of art; _i.e._ a product of one rational cause distinct
from the matter (of the parts), whose causality (in the collection and
combination of the parts) is determined through its Idea of a whole
possible by their means (and consequently not through external nature).
But if a thing as a natural product is to involve in itself and in its
internal possibility a reference to purposes,--_i.e._ to be possible
only as a natural purpose, and without the causality of the concepts of
rational beings external to itself,--then it is requisite _secondly_
that its parts should so combine in the unity of a whole that they are
reciprocally cause and effect of each other’s form. Only in this way
can the Idea of the whole conversely (reciprocally) determine the form
and combination of all the parts; not indeed as cause--for then it
would be an artificial product--but as the ground of cognition, for him
who is judging it, of the systematic unity and combination of all the
manifold contained in the given material.
For a body then which is to be judged in itself and its internal
possibility as a natural purpose, it is requisite that its parts
mutually depend upon each other both as to their form and their
combination, and so produce a whole by their own causality; while
conversely the concept of the whole may be regarded as its cause
according to a principle (in a being possessing a causality according
to concepts adequate to such a product). In this case then the
connexion of _effective causes_ may be judged as an _effect through
final causes_.
In such a product of nature every part not only exists _by means
of_ the other parts, but is thought as existing _for the sake of_
the others and the whole, that is as an (organic) instrument. Thus,
however, it might be an artificial instrument, and so might be
represented only as a purpose that is possible in general; but also
its parts are all organs reciprocally _producing_ each other. This can
never be the case with artificial instruments, but only with nature
which supplies all the material for instruments (even for those of
art). Only a product of such a kind can be called a _natural purpose_,
and this because it is an _organised_ and _self-organising being_.