During the long and peaceful reign of his son Angelo, little occurred to disturb the progress of the Duchy. But its external relations underwent a change at this time, in consequence of the transference of the suzerainty over it from the weak Emperor of Romania to the powerful Prince of Achaia, Geoffroy II, as a reward for Geoffroy’s assistance in defending the Latin Empire against the Greeks. Angelo, too, equipped three galleys for the defence of Constantinople, and, after its fall, sent a handsome present to the exiled Emperor. Like his father, he was summoned to aid the Venetian Governor of Crete against the native insurgents, but on the approach of the Nicene fleet he cautiously withdrew. His son, Marco II, who succeeded him in 1262, found himself face to face with a more difficult situation than that which had prevailed in the times of his father and grandfather. The Greeks had recovered ground not only at Constantinople, but in the south-east of the Morea, and their successes were repeated on a smaller scale in the Archipelago. Licario, the Byzantine admiral, captured many of the Ægean islands, some of which remained thenceforth part of the imperial dominions. Besides the Sanudi, the dynasty of the Ghisi, lords of Tenos and Mykonos, alone managed to hold its own against the Greek invasion; yet even the Ghisi suffered considerably from the attacks of the redoubtable admiral. One member of that family was fond of applying to himself the Ovidian line, “I am too big a man to be harmed by fortune,” and his subjects on the island of Skopelos, which has lately been notorious as the place of exile of Royalist politicians, used to boast that, even if the whole realm of Romania fell, they would escape destruction. But Licario, who knew that Skopelos lacked water, invested it during a hot summer, forced it to capitulate, and sent the haughty Ghisi in chains to Constantinople. Marco II had to quell an insurrection of the Greeks at Melos, who thought that the time had come for shaking off the Latin yoke. Educated at the court of Guillaume de Villehardouin, Marco had imbibed the resolute methods of that energetic prince, and he soon showed that he did not intend to relax his hold on what his grandfather had seized. Aided by a body of Frank fugitives from Constantinople, he reduced the rebels to submission, and pardoned all of them with the exception of a Greek priest whom he suspected of being the cause of the revolt. This man he is said to have ordered to be bound hand and foot, and then thrown into the harbour of Melos.