Why are the very early Paleozoic strata so rich in fossils, while the
immediately preceding Proterozoic rocks show so few? The seemingly
sudden appearance of so many highly developed animals in earliest
Paleozoic (Cambrian) time is one of the most important considerations
in the history of animal life, and it is by no means definitely
understood. The following statements bear directly upon the problem:
The early animal forms were probably soft or gelatinous without shells
and lived mostly in the open sea where food (seaweeds, etc.) was
abundant. Such animals were very unfavorable for preservation in fossil
form. Then, late in Proterozoic time or very early in the Paleozoic,
a severe struggle for existence set in, probably due to crowding
along shores, and hard parts began to develop both for support and
defensive purposes. Such hard parts or shells were commonly favorable
for fossilization. This view is strongly supported by the fact that
very thin shells only are known from late Proterozoic rocks, and mostly
very thin shells from the earliest Cambrian, the heavier shells having
been evolved later. A fact of importance to bear in mind in this
connection is that just at the critical time (late Proterozoic) in
shell development, the lands of the earth were undergoing widespread
and deep erosion as pointed out early in the chapter on "Ancient Earth
History." The earliest Cambrian strata, therefore, nearly everywhere
rest upon the deeply eroded surface of the Proterozoic rocks so that
the transition strata--the very ones which would contain most fossils
of the early shell development stage--are nearly everywhere missing.
Finally, mention should be made of the fact, that all Archeozoic
strata are profoundly altered (metamorphosed), and so are nearly
all Proterozoic strata, except the later. Fossils once present in
those rocks would of course have been obliterated by the process of
metamorphism, but the fact remains that very considerable thicknesses
of practically unaltered Proterozoic strata show few if any animal
fossils.
We shall now proceed to a rather systematic consideration of the most
interesting and significant types of creatures which have inhabited
the earth since the beginning of Paleozoic time at least twenty-five
million years ago. It is our purpose to bring out the salient features
in the history of each subkingdom of animals, beginning with the lowest
or simplest, and taking up in turn the higher and higher subkingdoms.
By this method the reader may easily follow the main thread of organic
evolution or progressive change which runs through most of the known
history of animal life of our planet, and which is so important in the
science of geology.