The Mollusks, which are more highly organized than the Molluscoids,
have more or less distinctly developed heads and locomotive organs.
Many thousands of species are now extinct, the classes of most
geological importance being represented by clams, snails, and the
pearly nautilus. Most of them have shells and gills for breathing. The
members of the simplest group, well represented by the clam tribe,
possess two similar shells working on hinges, so that in this regard
they are much like brachiopods, but, unlike the latter, they are not
symmetrical with reference to a plane at right angles to the hinge
line. Cambrian strata contain the oldest known of the fossil forms
where they are small, relatively thin-shelled, and rare. In marked
contrast to the brachiopods these bivalves have rather steadily
increased in numbers of species and individuals to the present time,
now being represented by thousands of forms. During the Mesozoic era
they greatly out-numbered the brachiopod bivalves and took on a more
distinctly modern aspect, when the oyster tribe and closely related
types were prominently developed. Culmination in size and thickness of
shell seem to have been reached in early Cenozoic time, strata of that
age in certain places, for example in Georgia and southern California,
being filled with oyster shells 10 to 20 inches long and 4 to 6 inches
thick! In addition to their gigantic size and thickness, many of the
shells were fluted or ribbed, and so they represented an extreme type
of defensive armor among the lower animals.
Snails have existed from the earliest Paleozoic era to the present
time, and the outstanding fact of interest concerning them is that
they furnish one of the finest illustrations of an important class
of animals which has undergone practically no conspicuous change or
evolution during all those millions of years of time.
[Illustration: Fig. 53.--Sketches of chambered cephalopods showing
the main steps in the evolution of the shell forms and compartment
partitions: a, b, the only kinds in Cambrian time; c, d, forms added
in the Ordovician; e, added in the Devonian; f, added in the late
Paleozoic; g, h, characteristic of the Mesozoic era; and i, a living
form (pearly nautilus) cut through. (Drawn by the author.)]