[Illustration: Fig. 58.--Restorations of characteristic Devonian
fishes, based upon actual fossils: a, a "lung fish" with leglike fins
(after Huasakof); b, a "ganoid." (After Nicholson.)]
In marked contrast to the most typical and highly organized fishes
so abundant to-day, all Devonian fishes were of simple types with
cartilaginous skeletons and vertebrated tails. Many of them were also
generalized types, that is, associated with their clearly defined fish
characters were others connecting them with certain higher Vertebrates,
as, for example, amphibians and reptiles. Thus all their tail fins
were vertebrated as in reptiles; their labyrinthine, internal tooth
structure was to be an amphibian feature when those creatures evolved;
many had protective armor or bony scales like most early amphibians and
many modern reptiles; and many had paired fins which were something
like jointed legs. Most abundant and highly organized of the Devonian
fishes were "ganoids," characterized by a covering of small plates or
bony scales set together but not overlapping like in typical modern
fishes. Their intricate tooth structure and limblike fins strongly
suggest the amphibians of later Paleozoic time. The skeleton of
cartilage gradually became somewhat ossified during succeeding geologic
periods. From their great profusion and diversity in the Devonian
period the ganoids have steadily fallen away until they now have very
few descendants like the gar pike.
Another important group of remarkable fishes, now totally extinct,
but common in Devonian and somewhat later time, had heavy, bony armor
plates over the fore part of the body. Those which grew to be fifteen
to twenty-five feet long were probably the rulers of the middle
Paleozoic seas. Another remarkable Devonian fish was able to breathe in
both water and air because, like their few modern descendants, they had
both gills and lungs. Because of their leglike fins and lung sac, it
is commonly believed that they were progenitors of the later Paleozoic
amphibians. The simplest of all fishes, the sharks, began in the
Silurian, underwent no important change through the millions of years
since, and are now of course well represented. During early Cenozoic
time the sharks seem to have reached culmination in size--sixty to
eighty feet long, with teeth five or six inches long.
Among modern fishes the most abundant by far, and the most highly
organized, are the true bony fishes, called the "teleosts," which
made their first appearance in the middle of the Mesozoic era. Those
earliest forms clearly show their descent from the ganoids. Apparently
they have not yet passed their prime.