The great diversity of modern placental animals may be suggested by
a few examples, as the tiger, dog, horse, camel, elephant, squirrel,
hedgehog, whale, monkey, and man. Forms like these, traced back
through their ancestors to the very early part of the Tertiary period,
gradually become less and less distinct until they cannot be at all
distinguished as separate groups, but rather there are ancestral
generalized forms which show combinations of features of the later
groups. Those early Tertiary generalized placental mammals had four
feet of primitive character, with five toes on each foot; the whole
foot, which from toe to heel touched the ground, was not adapted to
swift running; the teeth were simple (primitive) in type and of full
original number (forty-four); the toes were supplied with nails which
were about intermediate between real claws and hoofs in structure; and
the brain was relatively much smaller and simpler in structure than in
most modern mammals.
[Illustration: Fig. 66.--Chart showing the main features of interest
in the evolution of the horse family through several million years of
the present (Cenozoic) era of geologic time. (After Matthew, American
Museum of Natural History.)]
The history of the horse family furnishes an excellent illustration of
certain evolutionary changes among mammals. Skeletons of many species,
ranging from the early Tertiary to the present, have been found in
remarkable state of preservation representing every important change
in the history of the horse family. A study of the chart will make
clear some of the most striking changes which have taken place. The
oldest member of the horse family represented on the chart was about
the size of a small fox, with four toes and a degenerated fifth toe
(splint) on the front foot, and three toes and splint on the hind
foot. Since the chart was made a still more primitive form, even more
closely resembling the original five-toed ancestor, has been found.
Gradually the middle toe enlarged, while the others disappeared except
the two splints or very degenerate toes still left in the modern horse.
Increase in size of the animal and brain capacity accompanied these
changes. Also the teeth underwent notable change, and two originally
separate bones (radius and ulna) of the foreleg became consolidated
into a single stronger bone.
The even-toed hoofed mammals of to-day, like the deer, pig, and camel,
are also the product of evolution much like that of the horse, except
that two of the original five toes have been equally developed,
while the others have either greatly degenerated, as in the pig, or
disappeared entirely, as in the camel.