The Foot-prints of the Creator : $b or, The Asterolepis of Stromness — John Stuart Mill — John Shaqi
The Foot-prints of the Creator : $b or, The Asterolepis of Stromness
John Stuart Mill · en
But is not the “cartilaginous structure” of the Placoids analogous to
the embryonic state of vertebrated animals in general? Do not the other
placoid peculiarities to which the author of the “Vestiges” refers,—such
as the heterocercal or one-sided tail, the position of the mouth on the
under side of the head, and the rudimental state of the maxillaries
and intermaxillaries,—bear further analogies with the embryonic state
of the higher animals? And is not “embryonic progress the grand key to
the theory of development?” Let us examine this matter. “These are the
characters,” says this ingenious writer, “which, above all, I am chiefly
concerned in looking to; for they are features of embryonic progress, and
embryonic progress is the grand key to the theory of development.” Bold
assertion, certainly; but, then, assertion is not argument! The statement
is not a reason for the faith that is in the author of the “Vestiges,”
but simply an avowal of it; it is simply a confession, not a defence,
of the Lamarckian creed; and, instead of being admitted as embodying a
first principle, it must be put stringently to the question, in order to
determine whether it contain a principle at all.
In the first place, let us remark, that the cartilaginous structure
of the Placoids bears no very striking analogy to the cartilaginous
structure of the higher vertebrata in the embryonic state. In the case
of the _Delphinidæ_, with their soft skeletons, the analogy is greatly
more close. Bone consists of animal matter, chiefly gelatinous, hardened
by a diffusion of inorganic earth. In the bones of young and fœtal
mammalia, inhabitants of the land, the gelatinous prevails; in the old
and middle-aged there is a preponderance of the earth. Now, in the bones
of the dolphin there is comparatively little earth. The analogies of
its internal skeleton bear, not on the skeletons of its brethren the
mature full-grown mammals of the land, but on the skeletons of their
immature or fœtal offspring. But in the case of the true Placoids that
analogy is faint indeed. Their skeletons contain true bone;—the vertebral
joints of the Sharks and Rays possess each, as has been shown, an osseous
nucleus, which retains, when subjected to the heat of a common fire,
the complete form of the joint; and their cranial framework has its
surface always covered over with hard osseous points. But though their
skeletons possess thus their modicum of bone, unlike those of embryonic
birds or mammals, they contain, in what is properly their cartilage, no
gelatine. The analogy signally fails in the very point in which it has
been deemed specially to exist. The cartilage of the _Chondropterygii_
is a substance so essentially different from that of young or embryonic
birds and mammals, and so unique in the animal kingdom, that the heated
water in which the one readily dissolves has no effect whatever upon the
other. It is, however, a curious circumstance, exemplified in some of the