The Foot-prints of the Creator : $b or, The Asterolepis of Stromness
John Stuart Mill · en
That space comprised within the arch of the lower jaws, in which the
hyoid bone and branchiostegous rays of the osseous fishes occur, was
filled by a single plate of great size and strength, and of singular
form, (fig. 36;) and to this plate, existing as a steep ridge running
along the centre of the interior surface, and thickening into a massy
knob at the anterior termination, that nail-shaped organism, which I have
described as one of the most characteristic bones of the _Asterolepis_,
belonged. In the _Osteolepis_, the space corresponding to that occupied
by this hyoid plate was filled, as shown in fig. 14, by five plates of
not inelegant form; and the divisions of the arch resembled those of
a small Gothic window, in which the single central mullion parts into
two branches atop. In the _Holoptychius_ and _Glyptolepis_ there were
but two plates; for the central mullion, _i. e._ line of division,
did not branch atop; and in the _Asterolepis_, where there was no
line of division, the strong nail-like bone occupied the place of the
central mullion. The hyoidal armature of the latter fish was strongest
in the line in which the others were weakest. Each of the five hyoid
plates of the _Osteolepis_, or of the two plates of the _Glyptolepis_
or _Holoptychius_, had its own centre of ossification; and in the
single plate of _Asterolepis_, the centre of ossification, as shown by
the radiations of the fibre, was the nail-head. This head, placed in
immediate contact with the strong boxes of bone which composed the under
jaw, just where their central joining occurred, seems to have lent them
a considerable degree of support, which at such a juncture may have been
not unnecessary. In some of the nail-heads, belonging, it is probable, to
a different species of _Asterolepis_ from that in which the nail figured
in page 7, and the plate in the opposite page, occurred,—for its general
form is different, (fig. 37,)—there appear well-marked ligamentary
impressions closely resembling that little spongy pit in the head of the
human thigh-bone to which what is termed the round ligament is attached.
The entire hyoid-plate, viewed on its outer side, resembles in form the
hyoid-bone,—or cartilage rather,—of the spotted dog-fish, (_Scyllium
stellare_;) but its area was at least a hundred times more extensive
than in the largest _Scyllium_, and, like all the dermal plates of the
_Asterolepis_, it was thickly fretted by the characteristic tubercles.
In the Ray, as in the Sharks, the piece of thin cartilage of which this
plate seems the homologue, is a flat, semi-transparent disk; and there
is no part of the animal in which the progress of those bony molecules
which encrust the internal framework may be more distinctly traced, as if
in the act of creeping over what they cover, in slim threads or shooting
points,—and much resembling new ice creeping in a frosty evening over the
surface of a pool.
[Illustration: Fig. 37.
NAIL-LIKE BONE OF HYOID PLATE.