"The differences formerly supposed to exist between the grizzlies
and the big brown bears appear, in the light of the material now
available, to distinguish certain groups of species from certain
other groups, rather than the grizzlies collectively from the big
brown bears collectively. In other words, the differences between
the grizzlies on the one hand and the big brown bears on the other
are neither so great nor so constant as at one time believed. And
there are species which in the present state of knowledge cannot be
positively referred to either group. In fact, it seems at least
possible that certain species which appear to belong with the
grizzlies are closely related to certain other species which clearly
belong with the big brown bears. The typical brown bears differ from
the typical grizzlies in peculiarities of color, claws, skull, and
teeth. The color of the former is more uniform, with less of the
surface grizzling due to admixture of pale-tipped hairs; the claws
are shorter, more curved, darker, and scurfy instead of smooth; the
skull is more massive; the fourth lower premolar is conical, lacking
the sulcate heel of the true grizzlies. But these are average
differences, not one of which holds true throughout the group. Most
of the specimens in museums consist of skulls only, unaccompanied by
skins or claws, leaving a doubt as to the external characters; and
in old bears the important fourth lower premolar is likely to be so
worn that its original form cannot be made out. And, worst of all,
some of the grizzlies lack the distinctive type of premolar, leaving
only the skull as a guide to their affinities. The present
classification, therefore, must be regarded as tentative and subject
to revision....
"The present paper is merely a review of the existing state of
knowledge of the grizzlies and big brown bears of America and does
not include either the polar or the black bears. It is not intended
as a monographic revision, but aims to supply a list of the species,
together with descriptions and comparisons of adult skulls, chiefly
males. Little is said of external characters, for the reason that
little is known, only a few skins with claws being available for
study.
"LIST OF SPECIES AND SUBSPECIES OF GRIZZLY AND BIG BROWN BEARS, WITH
TYPE LOCALITIES.[1]
[Footnote 1: "Nearly 130 years ago Prof. Zauschner proposed the name
_Ursus saribur_ for an animal 'from the region of Canada' (Bestimmung
der Hundsart Krokute, und der Bärenart Saribur, p. 8, 1788), but the
species appears to be impossible of identification."]
(_Classification provisional._)