_Oaxaca_: 30 km. N of Matias Romero; Rio Sarabia (5);
Tapanatepec (87); Tolosita (6); between Zanatepec and
Tapanatepec. _Veracruz_: 25 km. SE of Jesus Carranza; 35 km.
SE of Jesus Carranza (2); 22 km. SSW of Jesus Carranza; 20
km. ENE of Jesus Carranza (7); Minatitlan; Tapalapan (5).
For the purposes of the present study I am not recognizing
_Eleutherodactylus beati_, _E. dorsoconcolor_, and _E. venustus_ as
specifically, or even subspecifically distinct from the earlier named
_E. rhodopis_. Probably these are mere color varieties of a single
species.
In the dry season frogs of this species were in humid forests, where
they were most frequently found along small streams and in ravines. The
species is widespread in the Gulf lowlands, but does not occur on the
Plains of Tehuantepec. It does inhabit the Pacific slopes on the
foothills of the Sierra Madre de Chiapas, the western part of which
extends into eastern Oaxaca near Tapanatepec.
=Eleutherodactylus rugulosus= Cope
_Oaxaca_: La Princesa (30); Modelo; Santa Lucia (10);
Tapanatepec (26); Tehuantepec (6); Tres Cruces (8).
_Veracruz_: Tapalapan (5).
In addition to the specimens from the lowlands of the isthmus, for the
purposes of the following discussion, I have included data on two
specimens from the southern slopes of the Sierra del Sur in Oaxaca
(Mirador and Chacalapa) and on several specimens from Los Tuxtlas in
Veracruz (Los Chaneques, 67; Salto de Eyipantla, 35; and San Andres
Tuxtla, 11).
Frogs of the _Eleutherodactylus rugulosus_ complex occur from southern
Veracruz and Sinaloa southward through Central America. Taylor
(1940:401) described _E. vocalis_ from Hacienda El Sabino, Michoacan;
Taylor and Smith (1945:580) described _E. avocalis_ from Tres Cruces,
Oaxaca. These have been considered as species distinct from
_rugulosus_, which is known to occur in Veracruz, Guerrero, and Chiapas
southward into Central America. Although the large number of specimens
collected in the isthmus does not aid in defining the ranges of the
taxa involved, these specimens do give some idea of the variation in
certain characters in a given population.
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