In specimens from Los Tuxtlas the tarsal fold is well-developed and
extends two-thirds to three-fourths the length of the tarsus; the
tibiotarsal articulation reaches the nostril and sometimes slightly
beyond the tip of the snout. In males the tympanum is nearly equal to
the diameter of the eye; in females it is about one-half the diameter
of the eye. The posterior surfaces of the thighs are dark brown or
black with whitish or cream-colored spots, flecks, or irregular
mottling. The tarsal fold is dark; the throat is pale in some
individuals, but in most is mottled with dark brown or gray flecks.
Individuals from La Princesa near the continental divide in Oaxaca show
the same variation in body proportions and development of the tarsal
fold. The posterior surfaces of the thighs are dark brown indistinctly
mottled with lighter brown. The throat is dark brown. Specimens from
the Pacific slopes of Oaxaca, including the Plains of Tehuantepec, have
dark brown thighs mottled with dusty cream. The tibiotarsal
articulation extends slightly beyond the tip of the snout in all
specimens. In males the tympanum is equal to about two-thirds the
diameter of the eye. Duellman (1958b:6) discussed the variation in
these characters in populations in Colima, Jalisco, and Michoacan.
Until the extent of variation of these characters is known throughout
the range of _rugulosus_, the recognition of populations either as
species or subspecies seems superfluous. Consequently, I have used the
oldest name; this does not necessarily imply, however, that all
populations of _rugulosus_ (_sensu lato_) are conspecific.
Of the 200 specimens examined, 15 have a middorsal stripe that is red
or yellow. The iris varies from a copper to a dark golden color and
shines bright red at night. Many of the specimens are juveniles; these
were collected in the dry season, at which time they were found beneath
rocks along streams, in road culverts where there was some water, and
in holes in banks and cliffs.
=Microbatrachylus pygmaeus= Taylor
_Oaxaca_: La Princesa (5); Matias Romero (9); Rio Sarabia
(41); Tolosita (2). _Veracruz_: Jesus Carranza; 20 km. ENE
of Jesus Carranza.
The specimens listed above vary widely in color patterns; some of the
patterns are characteristic of certain named "species": _albolabris_,
_imitator_, _lineatissimus_, and _minimus_. The large series from the
Rio Sarabia contains all of the color patterns; this series was
obtained in one small ravine in the rainforest. At least in the
isthmian region, this species does not inhabit the Pacific slopes and
lowlands.
=Syrrhophus leprus= Cope
_Oaxaca_: 33 km. N of Matias Romero; Santa Efigenia.
_Veracruz_: San Lorenzo.
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