A Guide to the Study of Fishes, Volume 1 (of 2)Jordan, David Starr
History
A Guide to the Study of Fishes, Volume 1 (of 2)
Jordan, David Starr
Fishes; Fishes -- Bibliography; Zoology -- History
In like fashion the spines and rays of the anal fin (18) are jointed
at base to bones called _interhæmals_ (77). In certain cases the
second interhæmal is much enlarged, made hollow and quill-shaped, and
in its concave upper end the tip of the air-bladder is received. This
structure is seen in the plume-fishes (_Calamus_). These two groups of
bones, interneural and interhæmal, are sometimes collectively called
_inter-spinals_. The flattened basal bone of the _caudal fin_ (80) is
known as _hypural_ (79).
[Illustration: FIG. 34.--Basal bone of dorsal fin, _Holoptychius
leptopterus_ (Agassiz). (After Woodward.)]
The tail of the striped bass, ending in a broad plate which supports
the caudal, is said to be homocercal. In more primitive forms the tail
is turned upward more or less, the fin being largely thrown to its
lower side. Such a tail as in the sturgeon is said to be heterocercal.
In the isocercal tail of the codfish and its relatives the vertebræ
are progressively smaller behind and the hypural plate is obsolete or
nearly so, the vertebræ remaining in the line of the axis of the body
and dividing the caudal fin equally. The simplest form of tail, called
diphycercal, is extended horizontally, tapering backward, the fin
equally divided above and below, without hypural plate. In any form of
the tail, it may through degeneration be attenuate or whip-like, a form
called leptocercal.
=The Pectoral Limb.=--The four limbs of the fish are represented by
the paired fins. The anterior limb is represented by the pectoral
fin and its basal elements with the shoulder-girdle, which in the
bony fishes reaches a higher degree of complexity than in any other
vertebrates. It is in connection with the shoulder-girdle that the
greatest confusion in names has occurred. This is due to an attempt
to homologize its parts with the shoulder-girdle (scapula, coracoid,
and clavicle) of higher vertebrates. But it is not evident that a
bony fish possesses a real scapula, coracoid, or even clavicle. The
parts of its shoulder-girdle are derived by one line of descent from
the undifferentiated elements of the cartilaginous shoulder-girdle of
ancestral crossopterygian or dipnoan forms. From a similar ancestry by
another line of differentiation has come the amphibian and reptilian
shoulder-girdle and its derivative, the girdle of birds and mammals.
=The Shoulder-girdle.=--In the higher fishes the uppermost bone of the
shoulder-girdle is called the _post-temporal_ (_suprascapula_) (53).
In the striped bass and in most fishes this bone is jointed to the
temporal region of the cranium. Sometimes, as in the trigger-fishes, it
is grown fast to the skull, but it usually rests lightly with the three
points of its upper end. In sharks and skates the shoulder-girdle,
which is formed of a continuous cartilage, does not touch the skull. In
the eels and their allies, it has, by degradation, lost its connection
and the post-temporal rests in the flesh behind the cranium.
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