The vegetative _branching_ is characteristic. One of the leaves
in a pair is formed before the other, and has a more vigorous
axillary bud; these stronger leaves stand in a ¼-spiral, the
fifth above the first one, and the branches are consequently
arranged in the same manner. In the inflorescence, however, it
is the upper or second bracteole (β) whose axillary bud (_w_ in
Fig. 361) is most advanced. The bud of the first bracteole (α)
becomes sometimes entirely suppressed, or in some this bracteole
itself is suppressed.
[Illustration: FIG. 360.--Diagram of _Lychnis_: α, β bracteoles.]
[Illustration: FIG. 361.--_A-F_ Diagrams of flowers of the
Caryophyllaceæ: _A Agrostemma_; _B Sagina_; _C Stellaria_;
_D Corrigiola_; _E Paronychia_; _F Herniaria_.]
The most original type appears to be represented by the Alsineæ.
From this form on one side the Sileneæ, adapted in a higher
degree for insect-pollination, are developed, and on the other
side the Paronychieæ, with various reductions.
=1.= ALSINEÆ, STITCHWORT GROUP. Sepals free, and connected with them
stellately expanded, slightly unguiculate (white or inodorous) petals;
these, however, often become suppressed (Fig. 363). The fruit is a
capsule.
[Illustration: FIG. 362.--_Cerastium arvense_: _A_ fruit; _B_
seed; _C_ section of seed.]
=a.= As many carpels as sepals (4 or 5). _Cerastium_ (Chickweed). The
petals are bifid. Capsule cylindrical, frequently _curved_ at the top,
and opening by 10 teeth (Fig. 362).--~_Malachium_ differs only in the
5-toothed capsule with bifid teeth.~--_Spergula_ (Spurry). The petals
are not bifid, capsule 5-valved; seeds winged. The leaves are linear,
and appear as if placed in large numbers in a whorl, a branch being
situated in the axil of each with leaves placed very close together at
its base; _stipules membranous_.--~Sagina has Sn, Pn, An + n, or An,
Gn, where n = 4 or 5. The corolla is often wanting.~
=b.= 3 (rarely 2) carpels (Fig. 361 _C_). _Stellaria_ (Stitchwort)
has deeply cleft petals. The number of stamens varies (see
above).--_Arenaria_ has entire petals. ~(To this group belong _Alsine_,
_Moehringia_, _Halianthus_, or _Honckenya_ (Fig. 363), which differ
from each other, especially in the form of the seed and number of
the capsular valves.) _Spergularia_ has membranous stipules, as in
_Spergula_.--_Holosteum._~
[Illustration: FIG. 363.--_Arenaria_ (_Halianthus_) _peploides_:
♀-(_A_) and ♂-flower (_B_, _C_).]
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