A history of land mammals in the western hemisphereScott, William Berryman
History
A history of land mammals in the western hemisphere
Scott, William Berryman
Mammals; Mammals, Fossil
So far as our information extends, there were no true Carnivora in the
Bridger, all the beasts of prey of the time belonging to the archaic
†Creodonta, which then reached their maximum development in numbers
and diversity. One family (†Oxyænidæ) included large and powerful
flesh-eaters, with cat-like dentition and short, rounded, lion-like
heads, long bodies and tails and short, heavy limbs, giving them the
proportions of otters. Another (the †Hyænodontidæ) comprised small,
long-headed, fox-like and weasel-like animals, which doubtless preyed
upon small mammals and birds. A third family (†Mesonychidæ) was made
up of moderate-sized, long-jawed creatures, which must have resembled,
rather remotely, short-legged and long-tailed wolves and hyenas. Their
habits and mode of life are somewhat problematical, for their grinding
teeth were blunt, not adapted to the shearing of flesh, and their claws
were broad, almost hoof-like. Such creatures could hardly have subsisted
by the pursuit of living prey and were probably carrion-feeders and more
or less omnivorous. The †Miacidæ, a family which connected the †creodonts
and true carnivores and might almost equally well be placed in either
group, were externally much like the small †hyænodonts, but were more
efficiently equipped for the capture and devouring of prey.
[Illustration: FIG. 139.—A mesonychid †creodont (_†Dromocyon velox_)
of the Bridger stage. Restored from a skeleton in the Museum of Yale
University.]
Of the archaic and extinct orders of hoofed animals, the only one which
persisted from earlier times into the Bridger and greatly flourished
there was the †Amblypoda, one family of which (†Uintatheriidæ) was
preëminently characteristic of middle Eocene life, becoming very rare
and then dying out in the upper Eocene. The †uintatheres of the Bridger
underwent considerable modification in size and appearance within the
limits of the stage, the larger and stranger species appearing toward
the end of the time. Most of these great creatures may fairly be called
gigantic, for they equalled the largest modern rhinoceroses and smaller
elephants in size. The body, limbs and feet were so elephantine in
character that they were once believed to be ancestral Proboscidea, but
the teeth and the fantastic skull were so radically different that this
belief was long ago abandoned. The upper canine teeth were converted, in
the males, into formidable spear-like or scimitar-like tusks, protected
by great flange-shaped expansions of the lower jaw; bony knobs on the
end of the nose probably supported a pair of dermal horns like those of a
rhinoceros and, in addition, a pair of high, cylindrical, horn-like, bony
protuberances arose above the eyes and another, more massive pair, near
the back of the head. It would be difficult to imagine more extraordinary
creatures than the †uintatheres, which were the largest land-mammals of
their time. The family was entirely confined to North America, no trace
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