A Mechanico-Physiological Theory of Organic EvolutionNägeli, Carl
Science
A Mechanico-Physiological Theory of Organic Evolution
Nägeli, Carl
Evolution
1. The essential part of the reproductive plasm, termed idioplasm, since
it divides and passes over from generation to generation, in higher as
well as in lower organisms, has a continuous or "immortal" existence.[I]
[I] Naegeli's idioplasm corresponds in many respects, though by
no means in all, to Weismann's germ-plasm. Weismann's idea of
continuity or "immortality," which has been so widely noticed,
is set forth with equal clearness, though with less emphasis, by
Naegeli.
2. During this continuous life the idioplasm goes through a development
of its own, just as an individual organism goes through a certain cycle
of development during its individual life. This development consists in
a constantly increasing complexity of structure and differentiation of
function.
3. This development is automatic, resulting from internal forces or
movements, (_Vervollkommnungs-bewegungen_).
4. As a result of the increasing complexity of structure in the
idioplasm the entire organism, which in each generation rearises
therefrom, becomes, from generation to generation, more and more complex
with greater and greater differentiation of function. Thus the
progression of the idioplasm controls the phylogeny of the race. It
marks out the course of evolution.
5. Since, according to Naegeli, new life with new idioplasms, may arise
wherever and whenever the necessary conditions combine, the present
organic world is not made up from branchings of a single original
idioplasm, but each race or group may have its own specific idioplasm;
and, since this has its own characteristic structure and its own
specific internal perfecting forces, it passes through its own peculiar
evolution, carrying with it its own depending race of organisms.
The fact that animals and plants at the present time show such various
degrees of organization is also accounted for on the last supposition,
for those of lowlier organization are merely of more recent origin and
have not progressed so far in idioplasmic development.
This automatic perfecting principle has been the mark of much criticism.
Some have confounded it with the mystical _nisus formativus_, or
formative principle of preceding theorists. But, as Weismann remarks,
Naegeli's phyletic force is conceived as a thoroughly scientific
mechanical principle. Naegeli has simply made application in the organic
world of the principle of entropy, as stated in the mechanical theory of
heat. Naegeli himself also compares his internal perfecting principle to
mechanical inertia. He says, "the force of evolution once started in a
given direction, tends to continue in the same direction. This
constitutes the law of inertia in the organic world."
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