A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
a special organ, occurring only in the Lepadidæ; the digestive organs
are very simple, from not having any distinct liver; the generative
system is very low, for both sexes are generally united in the same
individual; and the testes and ovaria closely resemble each other.
On the other hand, the thoracic limbs are, to a considerable extent,
specialised in their structure and functions; only the three posterior
pairs strictly resembling each other. Lastly, the dermal and muscular
systems are complicated, and not, to use Professor Owen's term, by mere
vegetative repetition, as will be obvious to any one who will study the
beautifully constructed and modified carapace--that is the operculum,
shell and basis--of a Balanus. On the whole, I look at a Cirripede
as a being of a low type, which has undergone much morphological
differentiation, and which has, in some few lines of structure, arrived
at considerable perfection,--meaning, by the terms perfection and
lowness, some vague resemblance to animals universally considered of a
higher rank.
[17] English Translation, in 'Scientific Memoirs,' 1853, vol. i, p.
228.
* * * * *
It has been seen that I divide the Cirripedia into three orders,--the
Thoracica, Abdominalia, and Apoda; between which the fundamental
difference consists in the limbs or cirri being thoracic in the first,
abdominal in the second; and entirely absent in the third. For the sake
of showing the range of character in Cirripedes, to which allusion has
been made, I will briefly indicate the leading differences in each
order. In the Thoracica, three families are included,--the Balanidæ,
or sessile Cirripedes, the Verrucidæ, remarkable from their quite
asymmetrical shell, and the Lepadidæ, or pedunculated Cirripedes. The
great difference in external appearance between these three families is
known to all naturalists. Even within the one family of Lepadidæ there
are great differences in external appearance, as will be admitted on
comparison of Lepas, Pollicipes, Conchoderma, &c.; but we have also
important internal differences, as in the case of Anelasma, in which
the cirri are barely articulated, and are not capable of seizing prey,
whilst the mouth is almost probosciformed, with the outer maxillæ
and palpi rudimentary: still more important are the differences in
Alcippe, in which the cirri of the first pair act as brushes; the
second, third, and fourth pairs being quite aborted; and the fifth and
sixth pairs consist only of four segments, with one of the two normal
rami converted into a crenated, button-like projection, for the sake
apparently of triturating food; Alcippe, also, is very remarkable in
being destitute of a rectum and anus. In this same genus Alcippe, in
Ibla and Scalpellum, there are either separate males or Complemental
males, some of which are so utterly abnormal in their characters, that
by no definition which I could frame, could they be included even in
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