A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
Podophthalmia; it leads into the Stomopoda, and likewise, as has been
noticed by many authors, into the sub-class Branchiopoda, which latter
sub-class is considered by Mr. Dana as only a part of the Entomostraca;
this group, therefore, exhibits affinities radiating in several
directions, and amongst these lines of relationship, one more must, I
believe, be added, plainly directed towards the Cirripedia.
[16] M. Martin St. Ange ('Mémoire sur l'Organ. des Cirripèdes,'
1835, extrait des 'Savans Etrangers,' tom. vi) has compared the
mouth of Lepas with that of Phyllosoma, and has given comparative
figures; but the resemblance is founded, I believe, on quite false
homologies.
One naturally wishes to ascertain how far Cirripedia are highly or
lowly organised and developed; but in all cases this, as it seems to
me, is a very obscure enquiry. Mr. Dana considers that, in Crustacea,
the greater or less centralisation of all the appendages round the
mouth is the main sign of high development; on this view, the anterior
part of a Cirripede, from being so much elongated, must be considered
as very low in the scale; the whole posterior part of the body, on the
other hand, is, in ordinary Cirripedes, brought close to the mouth; but
this is effected by the abortion of the seventh and eighth segments of
the cephalo-thorax and of the whole abdomen, and so, I presume, would
not, in Mr. Dana's estimation, raise the class much in the scale. Von
Baer[17] considers that the perfection of the type of any animal is in
relation to the amount of "morphological differentiation" which it has
undergone; on this view, Cirripedes ought to stand high in the scale,
for they differ much morphologically from the type of the class to
which they belong; as indeed is shown by the long time that elapsed
before their true position, namely amongst the Crustacea, was even
suspected; but something more must, I think, be added to Von Baer's
definition; for, to take as an example the eyes of a Cirripede,--as
seen in the first larval stage, there is only one eye, and that most
simple; in the pupa there are two, both compound, and furnished with
complicated muscles; lastly, in the mature animal there are still
two, but of very minute size, often almost confluent, and of the
simplest structure; hence, then, there has been much morphological
differentiation, but it is almost a contradiction in terms to speak, in
relation to such a case, of _perfection_ of type; and what has happened
to one organ, might happen to other organs, and so to the whole animal.
Lastly, under a physiological point of view, and taking the Balanidæ as
the most perfect type of the class, the sub-œsophageal portion of the
nervous system is highly concentrated; the organs of sense, excepting
the eyes, seem more largely developed than in ordinary Crustaceans; the
circulating system is of the simplest kind, being only lacunal; special
Branchiæ, however, are developed by the metamorphosis of, as I believe,
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