A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
In _Scalpellum Peronii_, and _villosum_, the males are but little
abnormal, for if classified independently of their sexual relations,
they would be considered as immature specimens of a new genus, standing
next to Scalpellum; in _Scalpellum rostratum_, the male would form
another and rather more distinct genus. The males, in the latter, are
attached to the other sex, between the basal edge of the labrum and
the adductor scutorum muscle; but in _S. Peronii_ and _villosum_ they
are attached lower down, in the furrow between the two scuta, and are
thus protected: in these three species, the internal parts of the male
present nothing particular. In Ibla, the males are attached low down
within the sack of the female; they may be said to consist of a mouth
surmounted on a long peduncle, for there is no capitulum or general
covering, and the whole thorax is in a rudimentary condition, the cirri
being reduced to two distorted pairs. As these males certainly moult
several times and grow a little, they must feed; and as they have no
cirri fit for action, they must seize their food by the contortions of
their peduncle, which we know homologically consists of the three first
segments of the head. The movements of the peduncle must, also, supply
those of the probosciformed penis, almost invariably present with other
Cirripedes, but here absent. If compelled to class these males without
regard to the female, great difficulty would be experienced; we could
hardly place in the family of the Lepadidæ, a Cirripede without a
capitulum, and without cirri, those very organs which give their name
to the class, and with a thorax reduced to the dimensions of a lower
lip; yet, if the presence of a peduncle did determine the classifier to
place these males amongst the Lepadidæ, then undoubtedly the character
of the mouth, &c. would fix their position next to Ibla.
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