Another very interesting feature of _Pelomyxa schiedti_ is the uroid
(Figure 6, _u_), which in this species consists of a number of very thin
projections resembling pseudopods extending from the posterior end.
These projections are attached to the substratum and in some way aid in
locomotion. These uroidal projections are of considerable length, and
may persist for a considerable length of time. Thus while _schiedti_ is
unable to form pseudopods at its anterior end, it forms uroidal
projections with great ease at its posterior end. But what the
conditions are which are necessary for the formation of a uroid, a
structure which it may be added, exists in many species of amebas (and
perhaps also in Cercomonas), is quite unknown.
In contrast to the amebas thus far discussed from the point of view of
the transformation of endoplasm into ectoplasm, there are a number of
species in which two distinct methods of endoplasmic transformation
occur typically. Among these species are the small _Amoeba radiosa_
(Figure 7), _A. bigemma_ (Figure 8) and a new species which for
convenience will be referred to as _bilzi_.
[Illustration: Figure 8. _Amoeba bigemma_, after Schaeffer. _a_, usual
form in locomotion, showing the numerous pseudopods, vacuoles, nucleus
and food body. _b_, rayed stage frequently assumed when suspended in the
water. The pseudopods in this stage are clear, slender, and more rigid
than those in stage _a_. _c_, an excretion sphere attached to a
twin-crystal characteristic of this ameba. _d_, the nucleus, consisting
of a clear nuclear membrane and a mass of chromatin granules in the
center. _e_, a small sphere attached to a crystal. _f_, a twin crystal
unattached to a sphere. Length of _a_, 150 microns; of _d_, 12 microns;
of _f_, 2 microns.]
It is well known that _radiosa_ has two stages: a more or less clavate
shaped stage in which the ameba creeps along the surface of some object
(Figure 7, _d_); and a stage in which a number (eight or less) of long
and very slender tapering pseudopods are formed which usually persist
for a long time (Figure 7, _a_, _b_). These pseudopods are frequently
quite straight and regularly disposed around the central mass of
protoplasm (Penard, ’02, pp. 87, 89). In no case are any endoplasmic
granules found in these slender pseudopods; they consist entirely of
hyaloplasm. In retracting these pseudopods a curious phenomenon is
sometimes observed; the pseudopod is rolled up into several (as many as
six) turns of an almost perfect helical spiral of a diameter six to
eight times that of the pseudopod. But as the process of withdrawal
proceeds, the spiral becomes irregular, but parts of some of the turns
persist in the last vestiges preceding complete withdrawal (Figure 7,
_b_). These spirals are also observed in other species besides _radiosa_
(see p. 128 seq.)
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