_Remarks._--As early as 1858 (p. 159) Baird recognized an individual of
this race from Pembina, Minnesota, as pertaining to a distinct species.
Although he used for it the specific name _pusillus_ originally
proposed by DeKay for a small weasel from the state of New York, Baird
wisely noted that the specimen he described "may be different from the
New York species. . . ." After preparing this account, Baird included a
second specimen, from Fort Steilacoom, Washington Territory, which he
thought might be the same, but the differences that he was careful to
point out, in the light of later knowledge, show it to be of the
species _Mustela erminea_. Only a few other naturalists followed Baird
in distinguishing the least weasel as a separate species until Bangs in
1896 (p. 21) clearly differentiated it and proposed for it the name
_Putorius rixosus_, which continues in use today and applies to the
species.
The accumulation at the National Museum of Canada, through the energy
of Dr. R. M. Anderson, of a good series of specimens from Saskatchewan
in the general vicinity of the type locality allows for the first time
an adequate conception of the amount of secondary sexual variation and
individual variation and permits recognition of subspecific characters
to differentiate between _M. r. rixosa_ and the subspecies _eskimo_ and
_campestris_. In comparison with the subspecies _allegheniensis_ the
basis for segregation is less clear and will remain somewhat in doubt
until additional adults of _allegheniensis_ from, say, Pennsylvania,
become available with accurate external measurements taken in the flesh
and especially with complete skulls.
Intergradation with the subspecies _eskimo_ is suggested by the short
tail of the specimen from fifteen miles east of Atlin, British
Columbia; in other particulars that specimen, a skin-alone, agrees with
the subspecies _rixosa_. Intergradation with _campestris_ is indicated
by increased size of some specimens from North Dakota, and is suggested
with _allegheniensis_ by the color of specimens from Wisconsin and
Illinois. Three specimens from Winona County, in southeastern
Minnesota, unfortunately are skulls-alone without external
measurements. Also, two of these skulls are of young animals. The one
adult, unsexed, is from Crystal Springs. Selected cranial measurements
are: basilar length, 28.5; length of tympanic bulla, 10.9. These
measurements accord with those of males of the subspecies _rixosa_ to
which the specimens from Winona County, therefore, are here assigned.
The possibilities have not been excluded, however, that the adult is an
unusually large female of the subspecies _campestris_ or a male of
_allegheniensis_ that has tympanic bullae longer than average for that
subspecies.
Public-domain text, read in full here on John Shaqi.
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