definitely for _frenata_. For convenience of handling, the population
is referred to _frenata_, providing that the average of specimens from
a nearby locality to the westward is not in favor of _arthuri_. In
event the average of specimens from a locality next adjacent to the
westward is in favor of _M. f. arthuri_, the total evidence from the
two localities may be weighed together and appropriate decision as to
subspecific status of weasels from the area is made according to what
the average is for the area as a whole.
The three individual animals of an intermediate sort are ordinarily
termed _intergrades_. This implies that their characters are the result
of mixed parentage--perhaps a female of _M. f. arthuri_ and a male of
_M. f. frenata_ but probably each parent itself was an intergrade and
the offspring, of which we examined three, owe their characters to
reproductive processes operating in obedience to Mendelian laws of
inheritance.
The two kinds of animals, _Mustela frenata arthuri_ and _Mustela
frenata frenata_, are identified as subspecies because of the
intergradation between them. If at this and all other places where the
geographic ranges of _arthuri_ and _frenata_ met there was no
crossbreeding (no intergrades), the two kinds would be treated as
distinct species. Intergradation, and the lack of it, are accepted as
the criteria of subspecies and species, respectively.
These criteria suffice for animals, in this instance weasels, which
have a continuous geographic distribution. Some kinds of weasels are
confined to islands, as for example the islands off the coast of Alaska
and British Columbia. Because weasels are land animals, crossbreeding
in nature between the weasels of two islands is, of course, impossible.
A modified test (used in the study here reported upon) in deciding on
specific versus subspecific status in these instances can be made as
follows: On the adjacent mainland, ascertain the degree of difference
between two subspecies whose geographic ranges meet (for example, _M.
e. richardsonii_ and _M. e. alascensis_). Next ascertain the degree of
difference between the insular kind of animal and the kind on the
mainland. If the degree of difference is greater when the insular kind
is compared than when only the kinds of the mainland are compared, the
insular kind is to be regarded as a species. If the degree of
difference is no greater between the insular kind and the mainland kind
than it is between the two adjacent mainland kinds, the insular kind is
to be regarded as a subspecies. In short, for insular kinds, the
criterion is degree of difference, with the limitation of geographic
adjacency, rather than intergradation.
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