The geographic variation (subspecific characters) found could be spoken
of as two kinds: First, there is the variation which is expressed in a
general trend for a long distance, producing, in general, a cline of
even slope; and second, that of inconstant trend in any one direction.
In his "The Rabbits of North America" Nelson (1909:34-35) has commented
on the latter type of variation as follows: "While studying series of
specimens from all parts of the vast range occupied by the geographic
races of such species as _Sylvilagus floridanus_ and _S. auduboni_, I
have been impressed with evidences of fluctuation of both external and
skull characters. These fluctuations are somewhat wavelike in character
and rise to central points of extreme development and then sink away to
intermediate borders beyond which new waves rise. Where the waves of
differentiation are pronounced they mark recognizable geographic races.
Within the area covered by the larger or geographically broader waves
of differentiation (recognized as of subspecific value), smaller waves
of differentiation are included, which may represent local variations
in intensity of characters of the subspecies, or these characters may
diminish and the variation tend in other directions, sometimes even
closely reproducing the characters of another subspecies occupying a
distinct area." In _Mustela frenata_, much of the geographic variation
at first inspection appears to be of this nature. Closer scrutiny,
however, reveals that the repetition, at geographic intervals, of
several features of color and structure are closely correlated with
environmental features which are repeated only at these same places.
In _Mustela erminea_, much of the variation is of the first kind,
namely, that which can be expressed as long clines of relatively even
slope. As several authors have said, zoölogical classification based on
this kind of variation is like dividing the spectrum and depends
largely upon the standards set, for, theoretically, the possibilities
of subdivision are unlimited. Actually, however, none of the clines has
an even slope and the possibilities for subdivision therefore are
limited. Also, when several features are used, instead of only one
feature, the classification is more satisfactory even if the basis is
more complex.
Some features of structure which provide subspecific characters are
mentioned below.
Total length, of males, ranges from 598 to 360 mm. in _M. frenata_ and
from 336 to 228 mm. in _M. erminea_. There is no cline of sustained
slope in _M. frenata_ but in _M. erminea_ there is a progressive
decrease in total length from north to south.
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