Length of tail varies from as little as a half to as much as
seven-tenths of the length of the head and body in _M. frenata_, the
subspecies _neomexicana_ having the long tail and the two subspecies
_arthuri_ and _primulina_ having short tails. The geographic ranges of
_primulina_ and _neomexicana_ are contiguous. In _M. erminea_ there is
likewise no variation of a clinal nature in length of tail and
furthermore the variation is much less than in _M. frenata_.
In length of hind foot, which in males varies from 49 mm. in northern
populations of _M. erminea_ to 28 mm. in southern populations, the same
cline is seen as in the total length of animals of this species. In _M.
frenata_, however, there are several decreases and increases along any
straight line which can be drawn through the geographic range of the
species. The range of variation in males is 41 mm. (_M. f.
arizonensis_) to 59 mm. (_M. f. macrophonius_).
Weight of the entire animal is an excellent measure of size but weights
are unavailable for many subspecies. In _M. frenata_, the two
subspecies _texensis_ and _macrophonius_ probably are the heaviest and
_effera_, _arizonensis_ and _helleri_ probably are the lightest.
Geographically the variation in weight behaves in approximately the
same way as does the measurement of total length. In _M. erminea_ the
variation in weight of males is from 206 grams in northern animals to
58 grams in southernmost populations, there being a relatively constant
gradient geographically.
Degree of hairiness of the foot-soles in _M. frenata_ clearly is linked
with the temperature; in regions of high average temperature the
hairiness is least and in regions of low average temperature it is
most. The decrease in hairiness is accomplished in two ways, namely,
smaller breadth and decreased length of individual hairs and decrease
in number of hairs on a given area of dermal surface. This correlation
holds throughout the entire north to south range of the species.
Corresponding differences are found on the same latitude where
topographic diversity in an east to west direction produces northern
conditions at high altitudes and southern conditions at low altitudes.
The conclusion seems unavoidable that climate, directly or indirectly,
determines the degree of hairiness. Less careful observations were made
on the hairiness of the soles of the feet in other species but it is
clear that the northern species _M. erminea_ has the most hair on the
foot-soles and that _M. africana_, the tropical weasel, has the least.
In this regard, _M. frenata_ is intermediate as it is also in
geographic position.
[Illustration: FIGS. 11-15. Dorsal views of adult skulls of each sex of
five subspecies of the ermine, _Mustela erminea_, to show secondary
sexual variation and geographic variation in size of the skull. Males
on the left and females on the right. All × 1.
Public-domain text, read in full here on John Shaqi.
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