An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
_Scelidotherium_, _Mylodon_, etc.—Of the more important remaining
genera of this family a briefer notice will suffice. _Scelidotherium_
(in which _Platyonyx_ may be included) comprises several species of
considerably smaller dimensions than the Megatherium, and is in some
respects intermediate between that genus and _Mylodon_. The teeth have
an oval cross-section, like those of the Sloths, while the skull, in
which the length of the nasals is subject to great variation in the
different species, approximates more or less closely to that of the
_Myrmecophagidæ_. The humerus generally has an entepicondylar foramen;
and the form and relations of the bones of the feet differ considerably
from those obtaining in the type genus. _S. leptocephalum_, the type of
the genus, occurs in Patagonia and Argentina but other species are found
in Brazil and Chili. The genus _Mylodon_, in its widest sense, may be
taken to include a number of comparatively large Edentates, some of which
have been described under the names of _Grypotherium_, _Lestodon_, and
_Pseudolestodon_. The teeth of the upper jaw are generally of an oval or
subtriangular section; and in the more typical forms the first and second
teeth are separated by a short interval, the former being horizontally
worn. In other species, however, like _M. (Lestodon) armatus_, there is
a considerable space between the first and second teeth, and the first
is worn obliquely. The skull is exceedingly like that of the Sloths in
general contour; and there is not the descending process at the angle of
the mandible found in _Megatherium_. The humerus has no entepicondylar
foramen. The species represented in Fig. 63 is from the Pleistocene of
South America; but the type of the genus is _M. harlani_, from beds
of corresponding age in Kentucky. The Patagonian _M. (Grypotherium)
darwini_ is a remarkable form, characterised by the presence of a
bony arch connecting the premaxillæ with the nasals, of which, as
already mentioned, there is an incomplete development in _Megatherium_.
_Megalonyx_, from the Pleistocene of Kentucky, differs from _Mylodon_ by
the long interval between the first and second teeth, and also by the
presence of an entepicondylar foramen in the humerus. _Nothrotherium_
is a smaller form, occurring in the deposits of the Brazilian caves, of
which the dental features have been already mentioned. The osteological
characters of these and other allied genera have been fully described
in the works of Cuvier, Owen, Burmeister, Leidy, Ameghino, Gervais,
Reinhardt, and others.
[Illustration: FIG. 63.—Skeleton of _Mylodon robustus_ (Pleistocene,
South America). From Owen.]
_Promegatherium._—Two genera from the infra-Pampean beds of Argentina,
described as _Promegatherium_ and _Promylodon_, are respectively
distinguished from _Megatherium_ and _Mylodon_ by the presence of bands
of enamel on the teeth, which points to the descent of the Edentates from
mammals with enamelled teeth.
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