When cross-sections of monocotyledonous and dicotyledonous bundles
are examined under the microscope, it is readily seen why
dicotyledonous bundles form rings of wood and monocotyledonous cannot
(Figs. 75 and 76). The dicotyledonous bundle (Fig. 76) has, running
across it, a layer of brick-shaped cells called =cambium=, which
cells are a specialized form of the parenchyma cells and retain the
power of growing and multiplying. The bundles containing cambium
are called _open bundles_. There is no cambium in monocotyledonous
bundles (Fig. 75) and the bundles are called _closed bundles_.
Monocotyledonous stems _soon cease to grow in diameter_. The stem
of a palm tree is almost as large at the top as at the base. As
dicotyledonous plants grow, the _stems become thicker each year_, for
the delicate active cambium layer forms new cells from early spring
until midsummer or autumn, adding to the wood within and to the bark
without. As the growth in spring is very rapid, the first wood-cells
formed are much larger than the last wood-cells formed by the slow
growth of the late season, and the spring wood is less dense and of
a lighter colour than the summer wood; hence the time between two
years’ growth is readily made out (Figs. 77 and 78). Because of the
rapid growth of the cambium in spring and its consequent soft walls
and fluid contents, the bark of trees “peels” readily at that season.
[Illustration: FIG. 77.—WHITE PINE STEM, 5 years old. The outermost
layer is bark.]
[Illustration: FIG. 78.—ARRANGEMENT OF TISSUES IN TWO-YEAR-OLD STEM
OF MOONSEED.
_p_, pith; _f_, parenchyma. The fibro-vascular bundles, or wood
strands, are very prominent, with thin medullary rays between.]
=Medullary Rays.=—The first year’s growth in dicotyledons forms a
woody ring which almost incloses the pith, and this is left as a
small cylinder which does not grow larger, even if the tree should
live a century. It is not quite inclosed, however, for the narrow
layers of soft cells separating the bundles remain between them (Fig.
78), forming radiating lines called =medullary rays= or =pith rays=.
[Illustration: FIG. 79.—MARKINGS IN CELL WALLS OF WOOD FIBRE
_sp_, spiral; _an_, annular; _sc_, scalariform.]
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