=The Several Plant Cells and their Functions.=—In the =wood= there
are some parenchyma cells that have thin walls still, but have lost
the power of division. They are now _storage cells_. There are also
wood fibres which are thick-walled and rigid (h, Fig. 76), and serve
to _support_ the =sap-canals= or _wood vessels_ (or tracheids)
that are formed by the absorption of the end walls of upright rows
of cells; the canals pass from the roots to the twigs and even to
ribs of the leaves and serve to transport the root water. They are
recognized (Fig. 79) by the peculiar thickening of the wall on
the inner surface of the tubes, occurring in the form of spirals.
Sometimes the whole wall is thickened except in spots called _pits_
(g, Fig. 76). These thin spots (Fig. 80) allow the sap to pass to
other cells or to neighbouring vessels.
[Illustration: FIG. 80.—PITS IN THE CELL WALL.
Longitudinal section of wall at _b_, showing pit borders at _o_,
_o_.]
The =cambium=, as we have seen, consists of cells whose function is
_growth_. These cells are thin-walled and filled with protoplasm.
During the growing season they are continually adding to the wood
within and the bark without; hence the layer moves outward as it
deposits the new woody layer within.
[Illustration: FIG. 81.—SIEVE-TUBES, _s_, _s_;
_p_ shows a top view of a sieve-plate, with a companion cell, _c_,
at the side; _o_ shows sieve-plates in the side of the cell. In
_s_, _s_ the protoplasm is shrunken from the walls by reagents.]
[Illustration: FIG. 82.—THICK-WALLED BAST CELLS.]
=The bark= consists of inner or _fibrous bark_ or new bast (these
fibres in flax become linen), the _green or middle bark_ which
functions somewhat as the leaves, and the _corky or outer bark_. The
common word “bark” is seen, therefore, not to represent a homogeneous
or simple structure, but rather a collection of several kinds of
tissue, all separating from the wood beneath by means of cambium. The
new bast contains (1) the _sieve-tubes_ (Fig. 81) which transport the
sap containing organic substances, as sugar and proteids, from the
leaves to the parts needing it (_s_, Fig. 76). These tubes have been
formed like the wood vessels, but they have sieve-plates to allow
the dense organic-laden sap to pass with sufficient readiness for
purposes of rapid distribution. (2) There are also thick-walled _bast
fibres_ (Fig. 82) in the bast that serve for _support_. (3) There is
also some parenchyma in the new bast; it is now in part a storage
_tissue_. Sometimes the walls of parenchyma cells in the cortex
thicken at the corners and form _brace cells_ (Fig. 83) (collenchyma)
for _support_; sometimes the whole wall is thickened, forming _grit
cells_ or _stone cells_ (Fig. 84; examples in tough parts of pear, or
in stone of fruits). Some parts serve for secretions (milk, rosin,
etc.) and are called _latex tubes_.
[Illustration: FIG. 83.—COLLENCHYMA IN WILD JEWELWEED OR TOUCH-ME-NOT
(IMPATIENS).]
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