How are they fixed to the skeleton? To see this all the smaller feathers and the muscles, or "flesh" of the wing must be removed. It will then be found that the flight-feathers are divisible into two series. One, widely spaced, runs along the upper surface of the fore-arm: the other, closely packed, along what answers to the back of the hand. In effect this is but a single rod of bone, but it is composed of three elements, answering to three of the digits of the human hand--the thumb and the first and second fingers. But they are scarcely recognizable as such, for the thumb is reduced to a mere stump, while the two fingers have become welded together. The third finger, indeed, has become reduced to the palm-bone, and a short stump answering to the first finger joint. To this frame-work, which can be folded up into the shape of a Z when the bird is at rest, the quills are fixed by their base by means of slender, but very strong elastic tendons. In birds which have a long upper arm bone, like the Albatross, Gull, or Heron, there is a third series of long, almost "quill-like" feathers running from the elbow to the body, thus closing up what would otherwise be a gap between the wing surface and the body, rendering flight impossible. [Illustration: Swans. Heron. Geese. ] The most important muscles of the wing are those which have to provide the power for the down-stroke of the wing. And these are the "pectoral" or "breast-muscles"--which form such dainty meat in a roast fowl. Owing to their great bulk the breast-bone itself would be insufficient to afford them attachment. This is furnished by the development of a deep, median keel, so that the breast-bone of a bird, such as a pigeon, bears a fanciful resemblance, when seen in profile, to the hull of a ship--unusually shallow--with a very deep keel. The front end of the breast-bone supports two slender rods of bone, and these in their turn support the long, sword-like blade-bone, and the "merry-thought." The general appearance of this frame-work for the support of the wing and its muscles can be seen in the adjoining illustrations. But it must be remembered that in their relative sizes and disposition these various parts present a very considerable range of differences. That these differences are correllated with different forms of flight goes without saying, but, be it noted, no one, as yet, has attempted to discover in what way they are related. Some of the readers of this book may, perhaps, be tempted to try and solve the problems which these differences present. To begin with, a collection of breast bones of different species of birds with their attached shoulder-girdles should be made, and these should be studied together with careful observations of the flight of the living bird. So far only a few comparisons of this kind have been made.