The discoveries of Garner and Allard as to the effect of light on reproduction began with a peculiar tobacco plant which appeared in some experiments at Washington. The plant grew to unusual size, and seemed to promise a valuable new variety. It formed no seeds, however, before the approach of cold weather. It was therefore removed to a greenhouse where it flowered and produced seed. In succeeding years the flowering was likewise delayed till early winter, but finally it was discovered that if small plants were started in the greenhouse in the early fall they flowered at the same time as the large ones. Experiments soon demonstrated that the time of flowering depends largely upon the length of the daily period when the plants are exposed to light. The same is true of many other plants, and there is great variety in the conditions which lead to flowering. Some plants, such as witch hazel, appear to be stimulated to bloom by very short days, while others, such as evening primrose, appear to require relatively long days. So sensitive are plants in this respect that Garner and Allard, by changing the length of the period of light, have caused a flowerbud in its early stages not only to stop developing but to return once more to a vegetative shoot. Common iris, which flowers in May and June, will not blossom under ordinary conditions when grown in the greenhouse in winter, even under the same temperature conditions that prevail in early summer. Again, one variety of soy beans will regularly begin to flower in June of each year, a second variety in July, and a third in August, when all are planted on the same date. There are no temperature differences during the summer months which could explain these differences in time of flowering; and, since "internal causes" alone cannot be accepted as furnishing a satisfactory explanation, some external factor other than temperature must be responsible. The ordinary varieties of cosmos regularly flower in the fall in northern latitudes if they are planted in the spring or summer. If grown in a warm greenhouse during the winter months the plants also flower readily, so that the cooler weather of fall is not a necessary condition. If successive plantings of cosmos are made in the greenhouse during the late winter and early spring months, maintaining a uniform temperature throughout, the plantings made after a certain date will fail to blossom promptly, but, on the contrary, will continue to grow till the following fall, thus flowering at the usual season for this species. This curious reversal of behavior with advance of the season cannot be attributed to change in temperature. Some other factor is responsible for the failure of cosmos to blossom during the summer months. In this respect the behavior of cosmos is just the opposite of that observed in iris.