Comments on the Taxonomy and Geographic Distribution of Some North American Marsupials, Insectivores and CarnivoresHall, E. Raymond (Eugene Raymond)
Science
Comments on the Taxonomy and Geographic Distribution of Some North American Marsupials, Insectivores and Carnivores
Hall, E. Raymond (Eugene Raymond)
Mammals -- North America
Before re-appraising the names it is pertinent to recall that Howell's
paper in 1906 on _Spilogale_ was only the second revisionary paper
that he prepared. It was prepared by a man who at that time lacked
much taxonomic experience, and who held to a morphotype concept.
Howell worked under the guidance, in the literal sense, of Dr. C. Hart
Merriam. The concept of species and subspecies held by Merriam
fortunately was recorded by him (Jour. Mamm., 1:6-9, November 28,
1919). Merriam's reliance on degree of difference and his disregard of
intergradation were naturally (and necessarily, we think, in Howell's
work in 1906) adopted by Howell. For example, of six specimens from
Point Reyes in west-central California, a place less than ten miles
from the type locality of _Spilogale phenax phenax_, Howell (_op.
cit._:33) assigned one specimen to the subspecies _Spilogale phenax
latifrons_! _S. p. latifrons_ occurs in Oregon and in northern
California--no nearer than 200 miles to Point Reyes. Howell's
assignment of this specimen to _S. p. latifrons_ was not a _lapsus_,
as persons with the modern (geographic) concept of a subspecies would
be likely to suppose. Howell's assignment of the one specimen to _S.
p. latifrons_ and the other five specimens to _S. p. phenax_ was
intentional, as he told one of us (Hall). He explained that he relied
upon the morphological characters of the individual animal instead of
upon the morphological characters of a population of animals. To him,
therefore, there was nothing inconsistent in his procedure in 1906.
Also, variation that was the result of difference in age and variation
that was the result of individual deviation were not understood, or at
least not taken into account, by Howell in 1906, nor by Merriam in
1890. For example, Merriam selected the most extensively white
specimen available to him for the holotype of _Spilogale leucoparia_.
He, and Howell in 1906, used the extensiveness of the white areas of
that particular specimen (see fig. 3, pl. 2, N. Amer. Fauna, 26, 1906)
as a character diagnostic of the "species" _S. leucoparia_ although
each of the authors had available two other specimens of _S.
leucoparia_ from the type locality, and all of the other referred
specimens in the United States National Museum, that were less
extensively white than the holotype. The _individual specimen_ was the
primary basis for the species or subspecies and one selected specimen
alone often was used in making comparisons between a given named kind
and some other species or subspecies. Also, be it remembered, degree
of difference, and not presence or absence of intergradation, was the
basis on which subspecific _versus_ specific rank was accorded to a
named kind of animal. Howell wrote on the labels of some specimens of
_Spilogale_ "not typical" when the individuals differed from the type
specimen in features that owe their existence to individual variation,
and he wrote the same words on the labels of other specimens that had
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