Cranial Osteology of the Hylid Frog, Smilisca baudiniTrueb, Linda
Science
Cranial Osteology of the Hylid Frog, Smilisca baudini
Trueb, Linda
Skull; Smilisca baudini
The bony dorsomedial part of the sphenethmoid diverges, forming the
anterior border of the frontoparietal fontanelle (_fpar. fon._, Figs.
1 and 14). The entire fontanelle is covered with a layer of dense
connective tissue continuous with that in which marginal bones and
cartilage of the sphenethmoid lie, and which is discrete from the
lower dermal layer of the overlying skin. At this level the braincase
is U-shaped in cross-section. Ossification terminates first in the
ventrolateral corners, followed by the lateral and dorsolateral areas.
The bony support of the latter area is furnished by the lamina
perpendicularis (_lam. perp._) of the frontoparietal (_fpar._, Fig.
14). Cartilage appears in the ventral part of the sphenethmoid in
posteromost sections; at the posterior levels of the orbit the
sphenethmoid is entirely cartilaginous.
Orbital, Otic, and Occipital Regions
_Orbital region._--The sclera (_scl._, Fig. 14) of the eye is
cartilaginous. The optic foramen (_opt. f._) is large and lies in
connective tissue at the posterior limits of the orbit and
sphenethmoid. At the posterior levels of the foramen the dorsolateral
cranial roof cartilages, taeniatecti marginales (_t. t. mar._)
converge medially to form the posterior margin of the frontoparietal
fontanelle and the tectum synoticum (_tect. syn._) of the occipital
region. At the posterior levels of the orbit the bursa angularis oris
(_b. ang. o._, Fig. 14) is present adjacent to the maxillary.
_Nerve foramina of otic and occipital regions._--The trochlear foramen
lies within the bony margins of the optic foramen. The trochlear nerve
is located posterodorsal to the optic tract, and separated from the
latter by connective tissue. The oculomotor foramen (_ocul. f._) lies
in connective tissue posterior and ventral to the optic foramen (Fig.
15). Anteriorly, dorsally, and ventrally the foramen has a bony margin
formed by the prootic (_pro._); posteriorly, only a thin layer of
connective tissue separates the oculomotor from the large prootic
foramen (_pro. f._). The latter is bordered by bone dorsally and by
cartilage ventrally (Fig. 16). Posteriorly, bone separates the prootic
foramen from the anterior acoustic foramen (_ant. acus._ _f._),
through which the ramus acusticus anterior and medius pass (Fig. 17).
An extremely narrow bridge of cartilage separates the anterior
acoustic foramen from the larger posterior acoustic foramen (_post.
acus. f._). The latter has a bony posterior margin and is widely
separated from the bony jugular foramen (_jug. f._) posteriorly
(Fig. 19).
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