Cranial Osteology of the Hylid Frog, Smilisca baudiniTrueb, Linda
Science
Cranial Osteology of the Hylid Frog, Smilisca baudini
Trueb, Linda
Skull; Smilisca baudini
_External dermal bones associated with the olfactory region._--The
association of the premaxillary (_pmax._) to the nasal cartilages is
described in preceding sections. The premaxillaries are separated from
each other medially and from the maxillaries laterally by dense
connective tissue. Anteriorly, the maxillary (_max._) bears a small
palatine process (_pal. proc._, Fig. 1) and a long, delicate pars
facialis (_p. fac._, Fig. 6), which terminates dorsally at the level
of the lamina inferior. Posterior to the transition zone between the
planum antorbitale and solum nasi the pars facialis is greatly
reduced. The pars palatina (_p. pal._, Fig. 6) persists to the
posterior part of the orbit.
The anterior end of the prevomer (_pvom._, Fig. 1) is associated with
the venter of the solum nasi at the level of the infundibulum just
posterior to the incorporation of the inferior prenasal cartilage into
the solum (Fig. 7). The prevomer expands dorsally around the distal
end of the solum to provide a bony lateral support for the olfactory
eminence (Figs. 10 and 11). A distal wing of the prevomer forms the
bony anterior and medial margins of the internal nares.
The palatine (_pal._, Figs. 1 and 12) lies in connective tissue medial
and adjacent to the pars facialis. At its maximum size the palatine
forms the bony posterior margin of the internal nares and extends
dorsomedially from the pars palatina to the distal part of the solum
nasi.
The nasal (_nas._, Fig. 1) is a thin bone overlying the tectum nasi
anteriorly (Fig. 7). It expands laterally to form a complete roof over
the cavum principale (Fig. 10). In the region of the internal nares,
the nasal forms the lateral wall of the cavum principale (Fig. 12).
Sphenethmoid Region
Posterior to the transition zone between the planum antorbitale and
solum nasi, the sphenethmoid (_spheth._, Fig. 1) is fully ossified
medially, the lateral parts of the bone at this level are only
ossified perichondrally. The septum nasi persists at the anterior
level of the orbit and terminates just anterior to the orbitonasal
foramen (_orbnas. f._) and the anterior end of the parasphenoid
(_pasph._, Fig. 13). The orbitonasal foramen is moderately large, has
a complete bony margin, and is located at the dorsolateral corner of
the braincase.
[Illustration: FIG. 13. Transverse section through sphenethmoid
region at level of orbitonasal foramen. Abbreviations: _ant. max.
proc._, anterior maxillary process; _max._, maxillary; _orbnas. f._,
orbitonasal foramen; _pasph._, parasphenoid; _spheth._, sphenethmoid.]
At the level of the orbitonasal foramen, the sphenethmoid is entirely
ossified except for a small dorsolateral extension. This distal
extension expands laterally in posterior sections as the braincase is
increased to its maximum width at the mid-length of the orbit; the
cartilaginous margin is retained throughout the length of the
sphenethmoid.
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