Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
of theories based on the fundamental postulate just mentioned--theories
having reference to the mechanism of heredity on the one hand, and to
the course of organic evolution on the other. Now it was the object of
the foregoing _Examination_ to deal with this system of theories _per
se_; and therefore we have here to take a new point of departure and to
consider separately the question of fact as to the inheritance or
non-inheritance of acquired characters. At first sight, no doubt, it
will appear that in adopting this method I am putting the cart before
the horse. For it may well appear that I ought first to have dealt with
the validity of Weismann's postulate, and not till then to have
considered the system of theories which he has raised upon it. But this
criticism is not likely to be urged by any one who is well acquainted
with the questions at issue. For, in the first place, it is notorious
that the question of fact is still open to question; and therefore it
ought to be considered separately, or apart from any theories which may
have been formed with regard to it. In the second place, our judgement
upon this question of fact must be largely influenced by the validity of
general reasonings, such as those put forward in the interests of rival
theories of heredity; and, as the theory of germ-plasm has been so
thoughtfully elaborated by Professor Weismann, I have sought to give it
the attention which it deserves as preliminary to our discussion of the
question of fact which now lies before us. Thirdly and lastly, even if
this question could be definitely answered by proving either that
acquired characters are inherited or that they are not, it would by no
means follow that Weismann's theory of heredity would be proved wholly
false in the one case, or wholly true in the other. That it need not be
wholly true, even were its fundamental postulate to be proved so, is
evident, because, although the fact might be taken to prove the theory
of Continuity, the theory of Germ-plasm is, as above stated, very much
more than this. That the theory of Germ-plasm need not be wholly false,
even if acquired characters should ever be proved heritable, a little
thought may easily show, because, in this event, the further question
would immediately arise as to the degrees and the comparative frequency
of such inheritance. For my own part, as stated in the _Examination_, I
have always been disposed to accept Mr. Galton's theory of Stirp in
preference to that of Germ-plasm on this very ground--i. e. that it does
not dogmatically exclude the possibility of an occasional inheritance of
acquired characters in faint though cumulative degrees. And whatever our
individual opinions may be touching the admissibility of such a _via
media_ between the theories of Pangenesis and Germ-plasm, at least we
may all agree on the desirability of fully considering the matter as a
preliminary to the discussion of the question of fact.
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