Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
As it is not to be expected that even those who may have read my
previous essay can now carry all these points in their memories, I will
here re-state them in a somewhat fuller form.
The following diagram will serve to give a clearer view of the sundry
parts of Professor Weismann's system of theories, as well as of their
relations to one another.
[Illustration: Postulate as to the absolute non-inheritance of acquired
characters.]
Now, as just explained, the parts of this system which may be properly
and distinctively called "Weismannism" are those which go to form the
Y-like structure of deductions from the fundamental postulate.
Therefore, it was the Y-like system of deductions which were dealt with
in the _Examination of Weismannism_, while it is only his basal
postulate which has to be dealt with in the following chapters.
So much, then, for the relations of Weismann's system of theories to one
another. It is, however, of even more importance that we should gain a
clear view of the relations between his theory of _heredity_ to those of
Darwin and of Galton, as preliminary to considering the fundamental
question of fact.
As we have already seen, the theory of germ-plasm is not only a theory
of heredity: it is also, and more distinctively, a theory of evolution,
&c. As a theory of heredity it is grounded on its author's fundamental
postulate--the _continuity_ of germ-plasm. But as a theory of evolution,
it requires for its support this additional postulate, that the
continuity of germ-plasm has been _absolute_ "since the first origin of
life." It is clear that this additional postulate is not needed for his
theory of heredity, but only for his additional theory of evolution, &c.
There have been one or two other theories of heredity, prior to this
one, which, like it, have been founded on the postulate of Continuity of
the substance of heredity; but it has not been needful for any of these
theories to postulate further that this substance has been _always_ thus
isolated, or even that it is now _invariably_ so. For even though the
isolation be frequently invaded by influences of body-changes on the
congenital characters of this substance, it does not follow that this
principle of Continuity may not still be true _in the main_, even
although it is supplemented in some degree by that of use-inheritance.
Indeed, so far as the phenomena of heredity are concerned, it is
conceivable that all congenital characters were originally acquired,
and afterwards became congenital on account of their long inheritance. I
do not myself advocate this view as biologically probable, but merely
state it as logically possible, and in order to show that, so far as the
phenomena of heredity are concerned, there appears to be no reason for
Weismann's deduction that the principle of Continuity, if true at all,
must be _absolute_. And it would further appear, the only reason why he
makes this deduction (stem of the Y) is in order to provide a foundation
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