Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
Now, although I allow that Darwin never attributed to this principle of
Amixia, or Independent Variability, anything like the degree of
importance to which, in the opinion of Delbœuf, Gulick, Giard, and
myself, it is entitled, the above passage appears to show that, as soon
as the "view" was clearly "suggested" to his mind, he was so far from
being unfavourably disposed towards it, that he added a paragraph to the
last edition of his _Variation_ for the express purpose of countenancing
it. Nevertheless, later on the matter appears to have entirely escaped
his memory; for in 1878 he wrote to Semper, that he did "not see at all
more clearly than I did before, from the numerous cases which he
[Wagner] has brought forward, how and why it is that a long isolated
form should almost always become slightly modified[41]." I think this
shows entire forgetfulness of the principle in question, because, if
the latter is good for explaining the _initial_ divergence of type as
between separated stocks of "domesticated animals," much more must it be
competent to explain the _further_ divergence of type which is "almost
always" observable in the case of "a long isolated form" under nature.
The very essence of the principle being that, when divergence of type
has once begun, this divergence must _ipso facto_ proceed at an
ever-accelerating pace, it is manifestly inconsistent to entertain the
principle as explaining the first commencement of divergence, and then
to ignore it as explaining the further progress of divergence. Hence, I
can only conclude that Darwin had forgotten this principle altogether
when he wrote his letter to Semper in 1878--owing, no doubt, as he says
in the sentence which immediately follows, to his having "not attended
much of late years to such questions."
[41] _Life and Letters_, vol. iii. p. 161.
* * * * *
So much, then, for Darwin's opinions. Next in order of time we must
consider Moritz Wagner's essays on what he called the "Law of
Migration[42]." The merit of these essays was, first, the firm expression
of opinion upon the swamping effects of free intercrossing; and, second,
the production of a large body of facts showing the importance of
geographical isolation in the prevention of these effects, and in the
consequent differentiation of specific types. On the other hand, the
defect of these essays was, first, not distinguishing between evolution
as monotypic and polytypic; and, second, not perceiving that
geographical isolation is only one among a number of other forms of
isolation. From these two radical oversights--which, however, were
shared by all other writers of the time, with the partial exception of
Darwin himself, as previously shown--there arose the following and most
lamentable errors.
[42] _Die Darwin'sche Theorie und das Migrationsgesetz_ (1868):
_Ueber den Einfluss der geographischen Isolirung_, &c. (1870).
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