Why varieties develop so readily and deviate so widely under domestication,
while they are apparently so rare or so transient in free Nature, may
easily be shown. In Nature, even with hermaphrodite plants, there is a vast
amount of cross-fertilization among various individuals of the same
species. The inevitable result of this (as was long ago explained in this
Journal [I-7]) is to repress variation, to keep the mass of a species
comparatively homogeneous over any area in which it abounds in individuals.
Starting from a suggestion of the late Mr. Knight, now so familiar, that
close interbreeding diminishes vigor and fertility; [I-8] and perceiving
that bisexuality is ever aimed at in Nature--being attained physiologically
in numerous cases where it is not structurally--Mr. Darwin has worked out
the subject in detail, and shown how general is the concurrence, either
habitual or occasional, of two hermaphrodite individuals in the
reproduction of their kind; and has drawn the philosophical inference that
probably no organic being self-fertilizes indefinitely; but that a cross
with another individual is occasionally--perhaps at very long
intervals--indispensable. We refer the reader to the section on the
intercrossing of individuals (pp. 96--101), and also to an article in the
Gardeners' Chronicle a year and a half ago, for the details of a very
interesting contribution to science, irrespective of theory. In
domestication, this intercrossing may be prevented; and in this prevention
lies the art of producing varieties. But "the art itself is Nature," since
the whole art consists in allowing the most universal of all natural
tendencies in organic things (inheritance) to operate uncontrolled by other
and obviously incidental tendencies. No new power, no artificial force, is
brought into play either by separating the stock of a desirable variety so
as to prevent mixture, or by selecting for breeders those individuals which
most largely partake of the peculiarities for which the breed is valued.
{I-9]
We see everywhere around us the remarkable results which Nature may be said
to have brought about under artificial selection and separation. Could she
accomplish similar results when left to herself? Variations might begin, we
know they do begin, in a wild state. But would any of them be preserved and
carried to an equal degree of deviation? Is there anything in Nature which
in the long-run may answer to artificial selection? Mr. Darwin thinks that
there is; and Natural Selection is the key-note of his discourse,
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