As a preliminary, he has a short chapter to show that there is variation in
Nature, and therefore something for natural selection to act upon. He
readily shows that such mere variations as may be directly referred to
physical conditions (like the depauperation of plants in a sterile soil, or
their dwarfing as they approach an Alpine summit, the thicker fur of an
animal from far northward, etc.), and also those individual differences
which we everywhere recognize but do not pretend to account for, are not
separable by any assignable line from more strongly-marked varieties;
likewise that there is no clear demarkation between the latter and
sub-species, or varieties of the highest grade (distinguished from species
not by any known inconstancy, but by the supposed lower importance of their
characteristics); nor between these and recognized species. "These
differences blend into each other in an insensible series, and the series
impresses the mind with an idea of an actual passage."
This gradation from species downward is well made out. To carry it one step
farther upward, our author presents in a strong light the differences which
prevail among naturalists as to what forms should be admitted to the rank
of species. Some genera (and these in some countries) give rise to far more
discrepancy than others; and it is concluded that the large or dominant
genera are usually the most variable. In a flora so small as the British,
182 plants, generally reckoned as varieties, have been ranked by some
botanists as species. Selecting the British genera which include the most
polymorphous forms, it appears that Babington's Flora gives them 251
species, Bentham's only 112, a difference of 139 doubtful forms. These are
nearly the extreme views, but they are the views of two most capable and
most experienced judges, in respect to one of the best-known floras of the
world. The fact is suggestive, that the best-known countries furnish the
greatest number of such doubtful cases. Illustrations of this kind may be
multiplied to a great extent. They make it plain that, whether species in
Nature are aboriginal and definite or not, our practical conclusions about
them, as embodied in systematic works, are not facts but judgments, and
largely fallible judgments-
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