Earthworms and Their AlliesBeddard, Frank E. (Frank Evers)
Science
Earthworms and Their Allies
Beddard, Frank E. (Frank Evers)
Earthworms
Finally, we have the largest of all the sub-families of this family,
viz. that of the Megascolecinae. These worms are mainly tropical in
range and also mainly found in the Indo-Australian portion of the
world. In them the sperm ducts open in common with the usually single
pair of spermiducal glands and prevalently upon the XVIIIth segment.
The glands moreover have not always, though they often have, the
tubular form shown in all the sub-families hitherto considered. In
many forms they are branched and lobate glands, and if there are two
pairs one may be of one type and the other of the second and derived
type, as for instance in _Megascolex ceylonicus_. Furthermore, it is
much commoner among the genera of this sub-family for the setae to
become numerous and to spread right round the segment; this condition
is seen in the genera _Pheretima_, _Megascolex_, _Diporochaeta_,
_Perionyx_, _Plionogaster_. The spermathecae also are commonly more
than the typical two pairs of the forms already considered, and in
certain species (for example _Pheretima hexatheca_) there are as many
as six pairs of those organs which are moreover--and in this they
resemble the majority of species of the last sub-families--nearly
always furnished with a diverticulum or diverticula. The nephridia
are either paired or numerous and these various characters allow of
the sub-family being split up into sixteen genera or thereabouts.
As an example of another type of organisation and as contrasting with
_Notiodrilus_ we may now briefly describe the structure of the genus
_Pontoscolex_ (better known as _Urochaeta_), a member of the American
and African family Geoscolecidae.
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