Earthworms and Their AlliesBeddard, Frank E. (Frank Evers)
Science
Earthworms and Their Allies
Beddard, Frank E. (Frank Evers)
Earthworms
Perhaps the next most widely distributed genus is _Helodrilus_ of
the family Lumbricidae which occupies Europe and Asia to the extreme
east, and is thought also to be indigenous to certain parts of North
America. But this range, though equally wide perhaps in mileage, is
less impressive than that of _Notiodrilus_, since the land areas
inhabited by the genus are continuous--almost so if we accept North
America as its real habitat. Here we have a case precisely the
opposite of that of _Notiodrilus_; for while there are reasons for
regarding _Notiodrilus_ as an ancient form of Lumbricid, there are
equally good reasons for regarding the Lumbricidae as the most modern
family of earthworms.
To find other instances of widely spread genera we must recur to the
great family Megascolecidae. There are practically no Geoscolecidae
which have a really extensive range. The only instances are
_Criodrilus_ and its ally _Sparganophilus_ which occur in America,
whether North or South, and in Europe; but as these forms are at
least largely aquatic the facts are not quite comparable with those
now under consideration.
The genus _Dichogaster_ (which includes as synonyms _Benhamia_,
_Millsonia_, _Microdrilus_) is unquestionably indigenous to tropical
Africa and certain parts of America including the West Indies. It has
been also met with in the East; but as the species there occurring,
such as for example the species originally described by myself as
_Microdrilus saliens_, are of small size, an accidental introduction
is quite possible, and it is by no means certain that it has not
occurred. In any case the genus is known to possess species which
are undoubtedly to be reckoned among peregrine forms--such as _D.
bolavi_, which has turned up in Europe. _Gordiodrilus_ and also
_Ocnerodrilus_ with its sub-genera have very much the same range as
has _Dichogaster_. It is to be noted however that these forms are
circumtropical, and that their distribution is thus less continuous
than that of _Notiodrilus_; they do not however show the markedly
discontinuous range of certain other genera of Megascolecidae. For
instance _Octochaetus_ is well known from New Zealand, and, not
occurring in the intermediate tracts, is again met with in India.
_Hoplochaetella_ is believed by Michaelsen to present us with another
precisely similar instance. Then also the genera _Woodwardia_ and
_Notoscolex_ are to be found in Australia and again (absent from the
immense tract lying in between) in Ceylon. _Megascolex_ has much the
same range, showing also this marked and remarkable discontinuity.
Stranger still, perhaps, is the range of _Plutellus_ and
_Megascolides_, of which the former, chiefly found in Australia and
Tasmania, not only extends its habitat to Ceylon but also to North
America; it is there represented by Eisen's species _Argilophilus
marmoratus_, referred by him, and not unnaturally, to a distinct
genus, but placed by Michaelsen in _Plutellus_. _Megascolides_ is
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