The shells require special study. In the lowest forms they are
membranous, sometimes encrusted with sand-grains, always very simple,
the only complication being the doubling of the pylome in _Diplophrys_
(fig. 2, 1), _Shepheardella_ (fig. 2, 3-5), _Amphitrema_ (fig. 2, 11),
_Diaphorophodon_ (fig. 2, 12). The marine shells are, as we have seen,
of cemented particles, or calcareous, glassy, and regularly perforated,
or again calcareous, but porcellanous and rarely perforate. These
characters have been used as a guide to classification; but some sandy
forms have so large a proportion of calcareous cement that they might
well be called encrusted calcareous genera, and are also not very
constant in respect of the character of perforation. The porcellanous
genera, however, form a compact group, the replacement of the shell by
silica in forms dwelling in the red clay of the ocean abysses, where
calcium carbonate is soluble, not really making any difficulty.
Moreover, the shells of this group show a deflected process or neck of
the embryonic chamber ("camptopyle") at least in the megalospheric
forms, whereas when such a neck exists in other groups it is straight.
The opening of the shell is called the pylome. This may be a mere hole
where the lateral walls of the body end, or there may be a diaphragmatic
ingrowth so as to narrow the entrance. It may be a simple rounded
opening, oblong or tri-multi-radiate, or branching (fig. 4, 1); or
replaced by a number of coarse pores ("ethmopyle") (fig. 3, 5a). Again,
it may lie at the end of a narrowed tube ("stylopyle"), which in
_Lagena_ (fig. 3, 9) may project outwards ("ectoselenial"), or inwards
("entoselenial"). In most groups the stylopyle is straight; but in the
majority of the porcellanous shells it is bent down on the side of the
shell, and constitutes the "flexopyle" of A. Kemna, which being a hybrid
term should be replaced by "camptopyle." The animal usually forms a
simple shell only after it has attained a certain size, and this
"embryonic chamber" cannot grow further. In _Spirillina_ and
_Ammodiscus_ there is no pylomic end-wall, and the shell continues to
grow as a spiral tube; in _Cornuspira_ (fig. 3, 1) there is a slight
constriction indicating the junction of a small embryonic chamber with a
camptopyle, but the rest of the shell is a simple flat spiral of several
turns. In the majority at least one chamber follows the first, with its
own pylome at the distal end. This second chamber may rest on the first,
so that the part on which it rests serves as a party-wall bounding the
front of the newer chamber as well as the back of the older; and this
state prevails for all added chambers in such cases. In the highest
vitreous shells, however, each chamber has its complete "proper wall";
while a "supplementary skeleton," a deposit of shelly matter, binds the
chambers together into a compact whole. In all cases the protoplasm from
the pylome may deposit additional matter on the outside of the shell, so
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