as to produce very characteristic sculpturing of the surface.
[Illustration: FIG. 3.--Various forms of Calcareous Foraminifera.
1, _Cornuspira_. 11, _Cristellaria_.
2, _Spiroloculina_. 12, _Globigerina_.
3, _Triloculina_. 13, _Polymorphina_.
4, _Biloculina_. 14, _Textularia_.
5, _Peneroplis_. 15, _Discorbina_.
6, _Orbiculina_ (cyclical). 16, _Polystomella_.
7, _Orbiculina_ (young). 17, _Planorbulina_.
8, _Orbiculina_ (spiral). 18, _Rotalia_.
9, _Lagena_. 19, _Nonionina_.
10, _Nodosaria_.]
Compound or "polythalamic" shells derive their general form largely from
the relations of successive chambers in size, shape and direction. This
is well shown in the porcellanous _Miliolidae_. If we call the straight
line uniting the two ends of a chamber the "polar axis," we find that
successive chambers have their pylomes at alternate poles; but they lie
on different meridians. In _Spiroloculina_ (fig. 3, 2) the divergence
between the meridians is 180 deg., and the chambers are strongly
incurved, so that the whole shell forms a flat spiral, of nearly
circular outline. In the majority, however, the chambers are crescentic
in section, their transverse prolongations being termed "alary"
outgrowths, so that successive chambers overlap; when under this
condition the angle of successive meridians is still 180 deg. we have
the form _Biloculina_ (fig. 3, 4), or with the alary extensions
completely enveloping, _Uniloculina_; when the angle is 120 deg. we have
_Triloculina_, or 144 deg., _Quinqueloculina_. Again in _Peneroplis_
(figs. 3, 5, and 4) the shell begins as a flattened shell which tends to
straighten out with further growth and additional chambers. In some
forms (_Spirolina_, fig. 22, 3) the chambers have a nearly circular
transverse section, and the adult shell is thus crozier-shaped. In
others (which may have the same sculpture, and are scarcely
distinguishable as species) the chambers are short and wide, drawn out
at right angles to the axis, but in the plane of the spiral, and the
growing shell becomes fan-shaped or "flabelliform" (figs. 3, 5, 4, 2).
This widening may go on till the outer chambers form the greater part of
a circle, as in _Orbiculina_ (fig. 3, 6-8) where, moreover, each large
chamber is subdivided by incomplete vertical bulkheads into a tier of
chamberlets; each chamberlet has a distinct pylomic pore opening to the
outside or to those of the next outer zone. In _Orbitolites_ (figs. 5,
6) we have a centre on a somewhat Milioline type; and after a few
chambers in spiral succession, complete circles of chambers are formed.
In the larger forms the new zones are of greater height, and horizontal
bulkheads divide the
chamberlets into vertical tiers, each with its own pylomic pore.
[Illustration: FIG. 4.--Modifications of _Peneroplis_. 1, _Dendritina_;
2, _Eu-Peneroplis_.]
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