We know that there was a time when the earth, hot and fiery, could
not have been the abode of life. Of the first origins of life we know
nothing and guess little. What we can justifiably surmise is that
the protoplasm of the original organisms was not yet differentiated
into cytoplasm and nucleus, and that sexuality had not yet arisen.
The bacteria, however specialized in other ways, are still in this
primitive condition.
Later, we can with great probability infer that the independent
units into which the stuff of life was subdivided reached a size
which, though still minute, was at least not beyond or even close to
the limits of microscopic vision; they were further provided with a
nucleus, and occasionally underwent sexual fusion. In other words, they
showed an organization which we call cellular; they were free-living
cells. Such unicellular creatures must have been at one epoch sole
inhabitants of the earth, and diverged into the most manifold types of
structure and modes of life. Such of them as led an animal as opposed
to a plant type of existence would be classified under the Protozoa or
unicellular animals.[5]
The colonial habit gives advantages of increased size and greater
rapidity of motion, of which many Protozoa have availed themselves. A
colonial existence once attained, division of labour, at first between
the germinal and the somatic, later between different types of somatic
units, will be a further advantage. Such organisms, of which we cannot
say definitely whether they are compound aggregates or single wholes,
would represent the most natural link between the unicellular Protozoan
and the rest of the animal kingdom, the multicellular forms or Metazoa.
And indeed such organisms exist at the present day--organisms such as
Volvox, Zoothamnium, Proterospongia, and Myxidium--as adjuvant and
confirmatory of our reasonable faith.
The multicellular organisms appear to have originated twice over, by
divergent routes. There are the true Metazoa, to which belong all the
higher types, and the Parazoa or sponges, which have never passed
beyond a very primitive type of structure. Both start as simple sacs,
whose walls are formed from two primary sheets or layers of cells.
Leaving sponges out of account, the Hydroid polyps are the simplest
representative of this grade of structure, while some of the Jelly-fish
and Siphonophores have attained the utmost limit of its inherent
possibilities.
The next great step was the intercalation of a third primary layer
between the other two. The result of this, the so-called triploblastic
type of organization, gives the ground-plan for all subsequent
organizations; and later evolution consists mainly in the evolution of
this ground-plan.
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