In other words, we can now pass from the consideration of the general plan of life’s architecture to that of its details. During the next great tract of time, that which was novel in life (for we must not be guilty of a _petitio principii_ in yet speaking of “advance” or “progress”) was brought about in two main ways--by an increase in the size of organisms, and by an increase in the efficiency of their working. The simplest Metazoa, such as the polyps, as well as the simplest three-layered forms, such as the free-living flat-worms, are all small, composed of an amount of material comparable with that contained in a single one of our hairs. In every group of Metazoa, increase of size is one of the main features that accompanies specialization, and the more specialized groups possess a higher average size than the less. A jelly-fish against a polyp; a cuttle-fish against a primitive mollusc; a vertebrate against its chordate ancestor; the giant reptiles of the late secondary period against their forbears; a horse against Phenacodus; man against the earliest primates--over and over again does size increase with the march of time. Not only this, but when there occurs aggregation of individuals to form units of a higher order, as in bees and ants and termites, and in man himself, there too increase of size in the new units thus produced is one of the most notable features. Is not human history in large measure the history of the increase in size of social units? But size alone is not enough; there is also a definite improvement of the details of life’s mechanism--partly revealed as improvement in the efficiency of the parts themselves, partly in the adjustment of the parts to each other, and their subordination to the needs of the whole. It is scarcely necessary to detail the improvements in efficiency of different organs during evolution: such are universally familiar. But a few examples will point my moral. The lowest three-layered forms have no circulatory system; this, rendered necessary later by increase of size, shows a gradual differentiation of parts in evolution. The exquisite machinery of our heart is directly descended from a minute pulsating ventral vessel such as that seen in Amphioxus. Protection and support are better cared for in insect than in worm, in mammal than in lamprey. But the most spectacular improvement of function, the most important of all the directional movements in evolution has been that affecting the nervous system and the sense-organs associated with it. Few people who have not gone carefully into the subject realize how imprisoned and windowless are the existences led by lower forms of life.