Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
Numerous specimens of geomyids from many localities and horizons are
available from the Pleistocene of North America. Specimens of the
genera _Geomys_ and _Thomomys_ are especially common. Few specimens
are known of the genera _Orthogeomys_ and _Pappogeomys_, especially
from the early and middle Pleistocene, owing, probably, to slight
knowledge of the early Pleistocene of México where these two genera
are thought to have evolved (see map, Figure 2). This lack of
knowledge about early Pleistocene deposits in México is a handicap in
the present instance since the center of differentiation for several
of the modern genera is judged to have been in México, probably on,
and at the edge of, the Central Plateau. The relative abundance of the
remains of _Geomys_ and _Thomomys_ from Pleistocene deposits farther
north, and the marked absence of other genera, may mean that
_Orthogeomys_ and _Pappogeomys_ did not range northward from southern
and central México in most of the Pleistocene. One species of
_Pappogeomys_ eventually ranged into the southwestern United States in
the late Pleistocene (toward the end of the Wisconsin) and it occurs
there today, but the genus is essentially Mexican.
The fossil record of _Zygogeomys_, as the genus is here understood,
evidently continued in the United States will into the Middle
Pleistocene, depending upon the stratigraphic interpretation of the
age of the Curtis Ranch local fauna from southeastern Arizona. Hibbard
(1958:25) regarded the Curtis Ranch local fauna as Irvingtonian in
age, a local fauna that lived either in the late Kansan glacial or the
Yarmouthian interglacial, and his correlation is tentatively followed
here. In deposits laid down later than those of Irvingtonian age no
remains of _Zygogeomys_ have been found. Today a single species exists
as a relic in the mountains of central México and _Zygogeomys_ may
have retreated southward to its present refugium in the late
Pleistocene. Perhaps, _Zygogeomys_ occurred in northern México and the
southwestern United States in the early and middle Pleistocene (see
Fig. 2), occupying the area between the ranges of _Pappogeomys_ to the
south and _Geomys_ to the north. Competition with _Pappogeomys_, and
especially _Geomys_, during Irvingtonian time may have extirpated
_Zygogeomys_ over most of this area, and by late Pleistocene
(Sangamon) much of the former range of _Zygogeomys_ came to be
occupied by one or the other of its competitors. The occurrence of
_Geomys garbanii_ in southern California (see White and Downs, 1961)
and the unidentified species of _Geomys_ in Aguascalientes (Mooser,
1959; for faunal correlation, see Hibbard and Mooser, 1963), both
from deposits of Irvingtonian age, supports this suggestion.
[Illustration: FIG. 2. Probable distribution of the Subfamily
Geomyinae in the early Pleistocene (late Blancan), depicting
major areas of differentiation of the modern genera.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account