Because fossils of complete fern plants have not yet been found,
separate names have been adopted for detached leaves, stems, and other
parts. For example, the fossil stems of some Pennsylvanian ferns found
in Illinois have been referred to two genera, _Megaphyton_, whose leaf
attachment scars are arranged in two vertical rows, one on either side
of the stem, and _Caulopteris_, whose leaf scars are arranged in a steep
spiral that becomes progressively flatter upward until near the top they
appear to be whorled. When the stem is a petrifaction, with internal
structures preserved, it is called _Psaronius_. The fronds are referred
to a number of genera, but those most commonly found in Illinois are
_Pecopteris_, _Asterotheca_, and _Ptychocarpus_.
[Illustration: Venation of seed fern leaflets]
_Pecopteris_
_Asterotheca_
_Ptychocarpus_
SEED FERNS
(Plate 4)
Seed ferns resembled true ferns in general, but they produced seeds,
borne on modified leaves. Where spore sacs and seeds are absent, the
leaves of seed ferns are difficult to distinguish from those of spore
ferns, although individual seed fern leaflets, called pinnae, are
somewhat larger.
[Illustration: _Medullosa_ Reconstruction and original drawing by
Wilson N. Stewart]
Seed ferns included vinelike plants in the undergrowth and trees such as
_Medullosa_. Some tree genera were very tall, with trunks more than two
feet in diameter. Unlike the true ferns, still living today, seed ferns
declined steadily after the close of the Pennsylvanian Period and
finally became extinct during Jurassic time. During Pennsylvanian time,
however, they were much more numerous and varied than true ferns.
[Illustration: Venation of seed fern leaflets]
_Alethopteris_
_Odontopteris_
_Mariopteris_
_Neuropteris_
_Linopteris_
Most of the common seed ferns found as fossils in Illinois can be
referred to the following leaf genera: _Alethopteris_, _Neuropteris_,
_Odontopteris_, _Linopteris_, _Mariopteris_ (which may be a true fern),
_Cyclopteris_, and _Spiropteris_. _Cyclopteris_ includes circular leaves
that occurred at the base of leaves referable to _Neuropteris_.
_Spiropteris_ includes young leaves that had not yet uncoiled and may
belong to either true ferns or seed ferns.
CORDAITES
(Plates 1 and 2)
[Illustration: Reconstruction of _Cordaites_
(after Hirmer)]
Cordaitean trees, forerunners of modern conifers such as pine and
spruce, were important during the Pennsylvanian Period for they were
distributed throughout the world. These trees, among the tallest plants
of the time, sometimes grew more than 100 feet high.
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